Calosoma (Callisthenes) anthrax Semenov, 1900

C. anthrax, represents the the easternmost species of Callisthenes. It is characterized by the pronotum with sharply protruding posterior lobes, by a rather squat shape and dark bronze color.
It is located in eastern Mongolia and in northern China.


Calosoma (Callisthenes) anthrax anthrax Semenov, 1900

Calosoma anthrax Semenov, 1900: 304 (type: au sud du lac of Buir nur, Zoological Museum of the Russian Academy of Sciences, Saint Petersburg)
Calosoma (Callisthenes) anthrax Breuning, 1928: 73
Callisthenes anthrax Lapouge, 1932: 376
Callisthenes anthrax Jeannel, 1940: 191

Length 21,5-25,5 mm. C. anthrax anthrax is characterized by transverse pronotum with conspicuously protruding posterior lobes. The sculpture of elytra consists of widespread raised graininess in which the foveae corresponding to the primary intervals are well evident. The color of the upper body is very dark with weak bronzed reflexes.
It has been described from the Mongolia's eastern border with China and later found in various localities of Northern China (Inner Mongolia and Shanxi).

Examined specimens and literature’s data
Mongolia: south of Buir nuur; 230km E SW of Choibalsan (Chaibalsan); Hulun nuur (Dalai-nor) (Obidov, 2002: 91)
China. Inner Mongolia: Hulun nuur (Dalai-nor) (Obidov, 2002: 91), Wuchuan (SB), Xilinhot (AC, SB); Shanxi: Dalong (GP).

Notes: Brachypterous. Adults were collected from mid May to August.

http://www.carabidae.ru/foto/thumbnails.php?album=1&page=1
Calosoma (Callisthenes) anthrax anthrax
Semenov, 1900
Chine: Inner Mongolia, Khing. south of Buir-Nur lake, 19.VI.1899, exp. Potanin (Zoological Institute, Russian Academy of Sciences, St. Petersburg) (Holotypus) (photo Anichtchenko A. 2010)
Calosoma (Callisthenes) anthrax anthrax
Semenov, 1900
Chine: Shanxi, 25 km north of Dalong, VII.2007, (coll. Pontuale).



Calosoma (Callisthenes) anthrax grumi Semenov, 1900

Calosoma Grumi Semenov, 1900: 306 (holotype: Danger-Chen, Zoological Museum of the Russian Academy of Sciences, Saint Petersburg)
Calosoma (Callisthenes) grumi Breuning, 1928: 74
Callisthenes anthrax grumi Lapouge, 1932: 376
Callisthenes grumi Jeannel, 1940: 191
Calosoma (Callisthenes) anthrax grumi Deuve 1997: 57

Length 22,5 mm. C. grumi was described by Semenov in 1900 on a single female specimen from East of Kuku-Nur in northern China (Qinghai), and never seen again. It slightly differs from C. anthrax because of the less developed lobes of the pronotum and because of the sculpture of elytra with more evident tegula and with the foveae, corresponding to primary intervals, reduced.
It was described as distinct species and as such kept by Breuning (1997: 57), Jeannel (1940: 191) and Obydov (2002: 90), but, according to Deuve (1997: 57), it is conspecific with C. anthrax, which opinion appears to be acceptable.

Examined specimens and literature’s data
China. Qinghai: Danger-Chenn (= Huangyuan) (Obydov, 2002: 91)

Notes: Brachypterous. The only known specimen was collected in late May.
The species was explicity dedicated by the author to the explorer G. Grum-Grzhimaylo or more exactly to Grigory Yefimovich Grumm-Grzhimaylo (1860–1936) a famous Russian entomologist and scientific traveller.

http://www.carabidae.ru/foto/thumbnails.php?album=1&page=1
Calosoma (Callisthenes) anthrax grumi
Semenov, 1900
Chine: Qinghai, Danger-Chen (Huangyuan) East of Kuku-Nor lake,
18.V. 1890, Grum-Grzhimailo (Zoological Institute,
Russian Academy of Sciences, St. Petersburg)
(Holotypus) (photo Anichtchenko A. 2010)


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