Calosoma (Callisthenes) ewersmanni Chaudoir, 1850

Callisthenes Ewersmanni Chaudoir, 1850: 157 (lectotype: Tokat; Muséum National d'Histoire Naturelle, Paris)
Calosoma persicum Gehin, 1885: 68 (holotype: Arménie Persane; Muséum National d'Histoire Naturelle, Paris)
Calosoma (Callisthenes) eversmanni, Breuning, 1928: 72
Calosoma (Callisthenes) eversmanni persicum, Breuning, 1928: 72
Callisthenes (Isocallia) breviuscula eversmanni, Lapouge, 1932: 373
Callisthenes panderi substriatus persicus, Lapouge, 1932: 375
Callisthenes eversmanni, Jeannel, 1940: 188
Callisthenes adehinotus Lassalle, 1992: 305 (holotype: Bingol, coll. Lassalle)
Calosoma (Callisthenes) pesarinii Heinz & Pavesi, 1994: 5 (holotype: Çorum, Mt. Elmali, coll. Heinz)
Calosoma (Callisthenes) peksi Heinz & Pavesi, 1994: 6 (holotype: Izmir, Boz-Dag, coll. Heinz)
Callisthenes eversmanni, Obydov, 2002: 84
Callisthenes eversmanni persicus, Obydov, 2002: 84
Callisthenes eversmanni pesarinii, Obydov 2002: 85
Callisthenes eversmanni adehinotus, Obydov, 2002,: 86
Callisthenes eversmanni peksi, Obydov, 2002: 86
Calosoma (Callisthenes) ewersmanni, Bousquet & al., 2003: 118
Callisthenes ewersmanni, Lorenz, 2005: 71


Length 19-27 mm. C. ewersmanni quite resembles to C. breviusculum but it is easily distinguishable because of its pronotum having only slightly elevated lateral margin with moderately pronounced basal dimples and with rear lobes broadly rounded, just protruding from the base line. It is also important to consider the sculpture of elytra that is characterized by coarse graininess, never so fine as in C. breviusculum, with primary intervals barely distinguishable by sparse foveae.
A moderate variability of C. ewersmanni has led to the description at species level of some of its populations: C. persicum (Gehin, 1885) with a golden green color and a stronger sculpture; C. pesarinii (Heinz & Pavesi, 1994) with greenish reflections on elytra and pronotum; C. peksi (Heinz & Pavesi, 1994) with less raised sculpture, C. adehinotus (Lassalle, 1992) with an even more flattened sculpture.
All the populations of what we consider the cluster of C. ewersmanni can be found in isolated stations of central northern Turkey, from Aegean coast (Izmir) up to Kurdistan (Lake Van), where its distribution area partially overlaps that of C. breviusculum. A possible penetration into western Iran (Ghahari & al 2009: 439) deserves confirmation, and the same applies to the presence of the species in the Syrian Anti-Lebanon mountains (http://carabidae.org/).

Examined specimens and literature’s data
Turkey. Izmir, Boz-Dag (holotype of peksi, Heinz & Pavesi, 1994: 6); Anatolien (SB); Tokat (type, NMHN); Çorum, Mt. Elmali (holotype of pesarinii, Heinz & Pavesi, 1994: 5), Samsun, Ankara, Eskisehir: Bos dagh (Breuning, 1928b: 73), Amasya (EM; SB), Tokat (holotype of Ewersmanni, MNHN), Merzifon (SB); Afiun-Karahissar (Afyon) (EM); Tuz Gölü, Aksaray, Sivas: Hafik (Heinz & Pavesi, 1994: 10); Bingol 1200m. (paratype of adehinotus, SB); Tunceli: Tunceli (NMP), Karaoglan (Heinz & Pavesi, 1994: 10); Elazığ: Yeniköy near Karakoçan 1200 m (sub persicum NMB), Dyarbakir (sub persicum Breuning, 1928b: 72).

Notes: Brachypterous. It lives on the Anatolia's plateau around 1200 - 1700m. altitude. Adults have been mostly collected in April - May.
The species was described as Callisthenes Ewersmanni because dedicated to the naturalist Alexander Eduard Friedrich Ewersmann (1794-1860) whose name, however, except for certain citations in Russian literature, it is commonly written Eversmann. The taxon has been therefore mostly erroneously cited as C. eversmanni.

Calosoma (Callisthenes) ewersmanni
Chaudoir, 1850
Tokat (lectotype of Calosoma ewersmanni Chaudoir, 1850)
(Muséum National d'Histoire Naturelle, Paris)
Calosoma (Callisthenes) ewersmanni
Chaudoir, 1850
Arménie Persane (holotype of Calosoma persicum Gehin, 1885)
(Muséum National d'Histoire Naturelle, Paris)
Calosoma (Callisthenes) ewersmanni
Chaudoir, 1850
Turkey: Anatolien (ex collection Najar, Négre)
Calosoma (Callisthenes) ewersmanni
Chaudoir, 1850
Turkey: Amasia, Asia min. sept.
Calosoma (Callisthenes) ewersmanni
Chaudoir, 1850
Turkey. Elazığ prov.: Yeniköy near Karakoçan, 1200 m, 19.IV.1981
(sub Calosoma (Callisthenes) eversmanni persicum Gehin, 1885)
(coll. Naturhistorischen Museum Basel)
Calosoma (Callisthenes) ewersmanni
Chaudoir, 1850
Turkey. Elazığ prov.: Yeniköy near Karakoçan, 1200 m, 19.IV.1981
(sub Calosoma (Callisthenes) eversmanni persicum Gehin, 1885)
(coll. Naturhistorischen Museum Basel)
Calosoma (Callisthenes) ewersmanni
Chaudoir, 1850
Turkey: Anatolie s.e, env. Bingol 1200m. V.1988,
Lassalle, (paratype Callisthenes adehinotus Lassalle, 1992)
(photo and coll. J. Matějíček)
Calosoma (Callisthenes) ewersmanni
Chaudoir, 1850
Turkey: Anatolie s.e, env. Bingol 1200m. V.1988,
Lassalle, (paratype Callisthenes adehinotus Lassalle, 1992)

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