Calosoma (Callisthenes) marginatum Gebler, 1830

C. marginatum and C. pseudocarabus were described as distinct species and as such considered, albeit doubtfully, by Jeannel (1940), followed in this by all subsequent authors. Both have small size and elongated Carabus type shape. Both are located in eastern Kazakhstan.
However, despite their geographical isolation and taking into consideration that they have many common characteristics and only slight morphological differences, are likely to be slightly different but conspecific populations.


Calosoma (Callisthenes) marginatum marginatum Gebler, 1830

Callisthenes marginatus Gebler, 1830: 307 (type: Nor-Saissan)
Calosoma (Callisthenes) panderi var. marginatum Breuning, 1928: 61
Callisthenes karelini marginatus Lapouge, 1932: 375
Callisthenes marginatus Jeannel, 1940: 190

Length 15-18 mm. You can sort out the typical population only by carefully examining the sculpture of elytra that is flat but slightly more evident than in C. pseudocarabus, and sometime observing greenish or bluish reflections on the edge of the pronotum and of the elytra, and on the bottom of the foveae of the primary intervals.
C. marginatum is located in the mountain ranges of Narym and Kurchum in south-eastern Kazakhstan.

Examined material
Kazakhstan: (N. of Zaisan lake) Narymsky range, Bolshenarymskoe (SB); Mt Kurchum, May-Terek (Obydov, 2002: 68).

Notes: Brachypterous. It lives at medium altitude around 1700. Adults were collected in June.

Calosoma (Callisthenes) marginatum marginatum
Gebler, 1830
S. Altai Mts, Narymsky Mt rng, Bolshenarymskoe env. 2100m, 12.VI.2000
Calosoma (Callisthenes) marginatum marginatum
Gebler, 1830
S.E. Kazakhstan: Narymsky range, N. Zaisan lake, 20.VI.2000.


Calosoma (Callisthenes) marginatum pseudocarabus Semenov-Tian-Shansky & Redikorzev, 1928

Calosoma pseudocarabus Semenov-Tian-Shansky et Redikorzev, 1928: 109 (type: Montis Saur a lacu Zaisan; Zoological Museum of the Zoological Institute of the Russian Academy of Sciences, Saint Petersburg)
Calosoma (Callisthenes) pseudocarabus Semenov, 1935: 274
Callisthenes pseudocarabus Jeannel, 1940: 190

Length 17-18 mm. C. pseudocarabus is slightly different from C. marginatum because its sculpture is more flattened, the foeae on the primary intervals are small, and in most cases barely visible, and because the color of the upper body is usually darker, rarely with faint blue reflections.
It is found on the Saur mountain range, also in south-eastern Kazakhstan.

Examined material
Kazakhstan: (S.E. of Zaisan lake) Saur mountain range, Mt Sargar (SB)

Notes: Brachypterous. It lives at medium altitude at 2000-2200m. Adults were collected from April to June.

Calosoma (Callisthenes) marginatum pseudocarabus
Semenov-Tian-Shansky & Redikorzev, 1928
S.E. Kazakhstan: Saur mts 2.000m., 12.IV.2000, Irtlach lgt.
Calosoma (Callisthenes) marginatum pseudocarabus
Semenov-Tian-Shansky & Redikorzev, 1928
S.E. Kazakhstan: Saur mts 2.000m., 12.IV.2000, Irtlach lgt.


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