Calosoma (Callisthenes) regelianum Morawitz, 1886

C. regelianum, as C. usgentense has elongated and convex elytra, with a smooth surface, where the striae are indicated by series of shallow punctures; the suture of elytra is more or less deeply depressed. However, C. regelianum is distinguishable by the proportionally larger head, the lateral margin of pronotum large, very strongly wrinkled. The margin of elytra is granulose, also in the basal region. The color of upper side of the body is generally glossy black. Mandl (1955: 328) described a species of northeastern Afghanistan, Calosoma (Callisthenes) klapperichi, that is very near to C. regelianum and that, at most, considering also the relative distance from other known populations, could be considered a subspecies of it. Consequently, the populations, related in some way to C. regelianum, occupy Tajikistan, and penetrate the north-eastern Afghanistan.


Calosoma (Callisthenes) regelianum regelianum Morawitz, 1886

Calosoma (Callisthenes) Regelianum Morawitz, 1886: 84 (lectotype: Baldshuan; Zoological Museum of the Zoological Institute of the Russian Academy of Sciences, Saint Petersburg)
Calosoma (Callisthenes) Regelianum var ovale Morawitz, 1886: 84 (described from: Baldshuan)
Calosoma (Callisthenes) Regelianum var oxygonum Morawitz, 1886: 84 (described from: Baldshuan)
Calosoma (Callisthenes) regelianum Breuning, 1928: 65
Callisthenes karelini regelianus Lapouge, 1932: 376
Callisthenes regelianus Jeannel, 1940: 193

Length 21-28 mm. C. regelianum regelianum is characterized by the pronotum rugged at the base and on the side and elytra enlarged in their rear half. The surface of elytra is almost smooth or with striae indicated by set of very shallow punctures. The color of the upper side of the body is entirely black or, rarely, with faint bronze luster on the margin of pronotum and on the border of elytra.
C. regelianum regelianum, is only found in Tajikistan.

Examined specimens and literature’s data
Tajikistan: eastern Bukhara (EM; SB); Tshitshantan (Breuning, 1928b: 66); Baldshuan (type, Breuning, 1928b: 66); Kulob, Garm valley (Breuning, 1928b: 66); khrebet Khozretishi 1600 m (SB); khrebet Petra Pervogo 2000 m (SB); Aksou valley, Kuljab (EM,SB), Darvas, 2500 m (EM); Tajikabad. 3000 m (SB)

Notes: Brachypterous. It is found mostly at altitudes around 1600-2000m., but it can also live up to 3500 m. Adults were collected in summer, from May to July.
Larval stages have been described by Mikhailov & Dzhabarova (1982).
The species is named in honour of Eduard August von Regel (1815–1892) a German botanist, director of the Botanical Garden of Saint Petersburg, who found the specimens object of the description

Calosoma (Callisthenes) regelianum
Morawitz, 1886
Tajikistan, Ganschou, Mt Pedra Pervogo,
vallis Sphedou 2000 m, 11-22.VI.1990, Relezar lg.
Calosoma (Callisthenes) regelianum
Morawitz, 1886
Tajikistan, Peter I rng. 3000m., 10km S Tajikabad
near Ganishod, 21.07.2003, O Pak leg.
Calosoma (Callisthenes) regelianum
Morawitz, 1886
Tajikistan, Ganschou, Mt Pedra Pervogo,
vallis Sphedou 2000 m, 11-22.VI.1990, Relezar lg.


Calosoma (Callisthenes) regelianum klapperichi Mandl, 1955

Calosoma (Callisthenes) klapperichi Mandl, 1955: 328 (holotype: Badakhshan, Shiva; ex coll. Klapperich)
Callisthenes sediquii Ledoux 1988: 156 (holotype: Badakhshan, Shiwa; coll. Ledoux)
Callisthenes klapperichi Obydov, 2002: 60

Length 20-25 mm. Mandl (1955: 328) described, at specific level, Calosoma (Callisthenes) klapperichi of northeastern Afghanistan. Obydov (2002) confirmed that it is a valid species and considered a synonym of it Callisthenes sediquii (Ledoux, 1988:156), described from the same place. We are speaking about a population, which, most probably, is part of the cluster of populations of C. regelianum, among which it stands out mainly because of the smaller and slender body with lesser developed head and slightly narrower elytra with almost parallel sides. The striae on the elytra are marked with clearly visible set of punctures that rarely are so evident in the individuals of the typical populations.
C. regelianum klapperichi has been only found in northern eastern Afghanistan. Instead in centre-western Afghanistan we find C. mirificum that is certainly an allied species, while in the centre-eastern part, according to the most recent findings, lives C. fabulosum, an ultra-advanced species, that could also be related to the same phyletic line.

Examined specimens and literature’s data
Afghanistan. Badakhshan: Shiwa (NMB); Panjshir: Ketal-el-Sehaba 3200 m (SB)

Notes: Brachypterous. It lives around 3200-3500 m. Adults were collected in June and July.
The taxon has been named after Johann Friedrich Klapperich (1913 - 1987) German entomologist and traveller, responsible for the Coleoptera at the Alexander-Koenig-Museum.

Calosoma (Callisthenes) regelianum klapperichi
Mandl, 1955
N.O. Afghanistan, Badakhshan, Shiwa Hochsteppe 2800m, 7.VII.73, Klapperich lgt.
(coll. Naturhistorischen Museum Basel)
Calosoma (Callisthenes) regelianum klapperichi
Mandl, 1955
N.O. Afghanistan, Badakhshan, Shiwa Hochsteppe 2800m, 7.VII.73, Klapperich lgt.
(coll. Naturhistorischen Museum Basel)
Calosoma (Callisthenes) regelianum klapperichi
Mandl, 1955
Afghanistan, Bad. Shiva, 14.VII.1954
(Obidov, http://www.zin.ru)

Calosoma (Callisthenes) regelianum klapperichi
Mandl, 1955
Afghanistan, Panjshir valley,
Ketal-el-Sehaba 3200m, VI.2002


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