Calosoma (Callitropa) anthracinum Dejean, 1831

C. anthracinum can be distinguished from the other apterous species previously collected in Blaptosoma for its more elongated body shape.
Jeannel (1940), followed in this by later authors, has considered the population of Guerrero State as the typical one ( anthracinum, including rufinum as a synonim) and interpreted as a subspecies (microgonum) all the other populations, that should differ from the first, mainly because of the barely visible basal dimples on pronotum, the average smaller size and even more slender shape.
At first, considering the low significance of these characteristics, we did not considered necessary the distinction between the two entities (Bruschi, 2013: 182).
Later Lassalle & van den Berghe (2013), have once more, taken into account the matter. They, having examined the male aedeagus of the types have clarified the synonimy of microgonum with rufinum and highlighted the differences of the two taxa with the typical anthracinum suggesting a distinction at species level between them, but we think that they should better be distinguished at the level of subspecies.
In the same publication, the authors described Calosoma (Blaptosoma) barksdalei (Lassalle & van den Berghe, 2013: 164) on two specimens that, from the description and from the images that accompany it, would seem to have some affinity with the populations of the anthracinum group. These specimens can be distinguished by the very slender and elongated shape similar to that of C. (Callitropa) anthracinum rufinum, but even more accentuated. However, always judging from the images, the apex of aedeagus seems similar to the one of C. (Callitropa) anthracinum anthracinum . For these reasons, at least for the moment we prefer to attribute these specimens to a further, distinct, population.


Calosoma (Callitropa) anthracinum anthracinum Dejean, 1831

Calosoma anthracinum Dejean, 1831: 569 (holotype: Mexico; Muséum National d'Histoire Naturelle, Paris)
Calosoma (Callitropa) laeve ab. anthracina Breuning, 1928: 46
) Blaptosoma (s. str.) laeve anthracinum Lapouge, 1932: 392
Callitropa (Blaptosoma) anthracinum Jeannel, 1940: 219
Calosoma (Blaptosoma) anthracinum anthracinum Gidaspow, 1959: 286
Calosoma (Blaptosoma) anthracinum (Lassalle & van den Berghe, 2013: 163)

Length 24-27 mm. The populations attributed to the typical form are characterized by the evident basal dimples of pronotum, the average larger size, the elytra slightly more rounded, the apex of aedeagus more stocky and obtuse.
All of these populations seem to be limited to an area between the state of Guerrero and the nearby state of Morelos.

Examined specimens and literature’s data
Mexico. (holotype of anthracinum, MNHN), Guerrero: Amula (= Almolonga) (AMNH); Morelos: Cuernavaca (Lassalle & Van Den Berghe, 2013:164).

Notes: Brachypterous. According to Erwin (2007: 87) it inhabits montane subtropical mixed oak-pine forest around 1800 m. Active adults have been captured in August.

Calosoma (Callitropa) anthracinum anthracinum
Dejean, 1831
Mexique (ex Musaeo Chaudoir) (holotype)
(coll. Muséum National d'Histoire Naturelle, Paris)
Calosoma (Callitropa) anthracinum anthracinum
Dejean, 1831
Guerrero, Amula 6000ft, Aug H H Smith 1891
(coll. American Museum of Natural History, New York)


Calosoma (Callitropa) anthracinum rufinum Géhin, 1885

Calosoma rufinum Géhin, 1885: 66 (holotype: Toluca; Muséum National d'Histoire Naturelle, Paris)
Calosoma laeve microgonum Bates, 1891: 225 (holotype: Jalapa, Natural History Museum, London)
Calosoma (Blaptosoma) laeve var microgonum Breuning, 1928: 46
Blaptosoma (s. str.) laeve microgonum Lapouge, 1932: 392
Callitropa (Blaptosoma) anthracinum microgonum Jeannel, 1940: 219
Calosoma (Blaptosoma) anthracinum microgonum Gidaspow, 1959: 286
Calosoma (Blaptosoma) rufinum (Lassalle & van den Berghe, 2013: 164)

Length 22-25 mm. C. anthracinum rufinum is distinguished by the basal dimples of pronoto hardly notable, the average smaller size, the more slender body shape, the apex of aedeagus relatively more elongated and slender.
C. anthracinum rufinum has been sporadically found in various States of central Mexico: Mexico, Puebla, Vera Cruz.

Examined specimens and literature’s data
Mexico. Jalisco: Chapala (Gidaspow, 1959: 286); Mexico: Toluca (holotype of rufinum, MNHN); Puebla: Esperanza (AMNH); Vera Cruz: Jalapa (EM).

Notes: Brachypterous. It lives in the Mexican plateau between 1500 and 2500m.

Calosoma (Callitropa) anthracinum rufinum
Géhin, 1885
Mexique, Toluca (holotype of Calosoma rufinum Géhin)
(coll.Muséum National d'Histoire Naturelle, Paris)
Calosoma (Callitropa) anthracinum rufinum
Géhin, 1885
Mexico, Vera Cruz (Jalapa) (coll. E. Migliaccio)


Calosoma (Callitropa) anthracinum barksdalei Lassalle & van den Berghe, 2013

Calosoma (Blaptosoma) barksdalei Lassalle & van den Berghe, 2013: 164 (holotype: nord de Guadalajara, coll. Lassalle)

Length 22-23 mm. As C. anthracinum rufinum it has an elongated body shape, with hardly notable basal dimples of pronotum and reduced hind angles, not extending beyond the basal line, but it can be differentiated by its just transverse pronotum, the elytra distinctly more elongated, the stouter apex of aedeagus.

Examined specimens and literature’s data
Only the images accompanying the original description.

Notes: Brachypterous. The only known two specimens, males, were collected north of Guadalajara. It is possible that the quotation of Gidaspow (1959: 286) of Calosoma (Blaptosoma) anthracinum microgonum (= Calosoma (Callitropa) anthracinum rufinum) from Chapala, about 50 km. south of Guadalajara, can be referred to this new subspecies that would be so localized in the central area of state of Jalisco.

Calosoma (Callitropa) anthracinum barksdalei
Lassalle & van den Berghe, 2013
holotype: nord de Guadalajara, coll. Lassalle
Lassalle & van den Berghe, 2013: 165, pl. 8

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