Calosoma (Callitropa) atrovirens Chaudoir, 1869

Length 22-28 mm. C. atrovirens, as other species previously collected in Blaptosoma, is characterized by the squat and convex body shape. In particular, C. atrovirens shares with C. laeve, a very bulky form and a strongly transverse pronotum, but it can be easily identified taking into account the presence of large dots on the head, from the vertex towards the eyes.
Jeannel (1940:218) had distinguished three subspecies (atrovirens, obscurum and explanaticolle). Gidaspow (1959: 282) considered superfluous explanaticolle, which should have differ from obscurum because of more elongated, and by more arched sides of pronotum, and she clarified the geographical distribution of species.
Most of subsequent authors agreed on the identity of the two supposed subspecies, but the name obscurum, that for priority reason could not be used, was replaced by explanaticolle.
The populations that would be considered typical are the oriental ones that are characterized by slender body and by the dark green color of the upper body, while those attributable to explanaticolle would be limited to the central western part of the Sierra Madre Occidental, and are distinguished from the typical ones by the black color, at most faintly greenish on body margins, and by the more squat shape.
But it happens that the only certainty is that the color varies from the eastern populations to the western ones, from dark green, spread over the entire upper body, to black or reddish black with faint greenish reflections at the margins of pronotum and elytra. The body shape instead varies greatly inside a same population and can not be used effectively as a diagnostic character.
However, considering that, as noted by Jeannel (1940: 218), the progressive loss of color is, in this case, the indicator of an evolutionary process in progress, we believe for the purposes of this discussion that these populations could be dealt with at subspecific level.


Calosoma (Callitropa) atrovirens atrovirens Chaudoir, 1869

Calosoma atrovirens Chaudoir, 1869: 372 (lectotype: interieur du Mexique; Muséum National d'Histoire Naturelle, Paris)
Calosoma laeve ab. atrovirens Breuning, 1928: 44
Blaptosoma (s. str.) laeve atrovirens Lapouge, 1932: 393
Callitropa (Blaptosoma) atrovirens Jeannel, 1940: 217
Callitropa (Blaptosoma) atrovirens Gidaspow, 1959: 282
Callitropa (Blaptosoma) atrovirens Erwin, 2007: 88

Length 25-30 mm. C. atrovirens atrovirens is characterized by the dark green colour of the upper side of the body and, on average, by a more elongated shape.
The label of the lectotype shows no locality details and the author in the description gives generically “interieur du Mexique” as the origin of the specimen. Most of the known specimens were found in Hidalgo. However, according to more recent data, C. atrovirens atrovirens has a larger distribution area and the populations that would be considered typical are the oriental ones (Eastern Guanajuato, Hidalgo Puebla, Queretaro, Tapaulimas (Ciudad del Maiz, on the border between San Luis Potosi and Tapaulimas).

Examined specimens and literature’s data
Mexico. interieur du Mexique (lectotype MNHN); Guanajuato: 10 km NW of Leon (UASM), 58 km S of San Luis de La Paz (UASM), Hidalgo: Tula (SB); Mexico: Teotihuacan (UASM); Michoacán: nearby Moralia (EM, GP, SB), Zitacuaro (UASM), Sahuayo de Morelos (Lassalle & van den Berghe, 2013:163), Jiquilpan (AMNH); Querétaro: Landa de Matamoros (UASM); Puebla: Tlaxaca (UASM); Tapaulimas: NW of Ciudad del Maiz (Gidaspow, 1959: 282).

Notes: Brachypterous. Lives on oak forests on midlands between 1100 and 1300m. Active adults have been captured in July-August and November (Erwin, 2007: 88)

Calosoma (Callitropa) atrovirens atrovirens
Chaudoir, 1869
interieur du Mexique (lectotype)
(coll. Muséum National d'Histoire Naturelle, Paris)
Calosoma (Callitropa) atrovirens atrovirens
Chaudoir, 1869
Hidalgo, Tula


Calosoma (Callitropa) atrovirens explanaticolle Bates, 1891

Calosoma obscurum Géhin, 1885: 65 (type: Oaxaca; Muséum National d'Histoire Naturelle, Paris) (preoccupied by Calosoma auropunctatum var. obscurum Letzner, 1850)
Calosoma explanaticolle Bates, 1891: 225
Calosoma laeve ab. atrovirens var. obscurum Breuning, 1928: 44
Calosoma laeve explanaticolle Breuning, 1928: 44
Blaptosoma (s. str.) laeve explanaticolle Lapouge, 1932: 393
Callitropa (Blaptosoma) atrovirens explanaticolle Jeannel, 1940: 217
Callitropa (Blaptosoma) atrovirens obscurum Jeannel, 1940: 217
Callitropa (Blaptosoma) atrovirens obscurum Gidaspow, 1959: 282
Callitropa (Blaptosoma) atrovirens explanaticolle Erwin, 2007: 88
Calosoma (Blaptosoma) atrovirens obscuroides Lassalle & van den Berghe, 2013: 163 (nomen novum pro obscurum Géhin, 1885).

Length 22-28 mm. Unlike the populations attributable to the typical form, those of C. atrovirens explanaticolle have the upper side of the body always devoid of metallic lustre showing a black or reddish black colour. These populations are found in some States of the Sierra Madre Occidental (Aguascalientes, Durango, Western Guanajuato , Jalisco, Michoacan, Zacatecas).
Recently, Lassalle & van den Berghe (2013: 163), came back to the distinction between obscurum and explanaticolle and, based on the morphological characteristics of the holotype of C. obscurum, especially the considerable width of the pronotum and the stronger marks on the head, they have reassessed it at subspecies level and renamed obscuroides. The holotype is the only known specimen and its provenance from Oaxaca is dubious.

Examined specimens and literature’s data
Mexico. Aguascalientes: around the city (Lassalle & van den Berghe, 2013:163); Durango: Alemán, reserva La Michilia (sub atrovirens, Ball & Shpeley, 1991: 43); Guanajuato: around Guanajuato city, Jerecuaro (Lassalle & van den Berghe, 2013:163), Tupataro (Breuning, 1928b: 46); Jalisco: Lagos de Moreno (UASM),Tepatitlan (AMNH), Chapala (Gidaspow, 1959: 283); Oaxaca (holotype of obscurum, MNHN); Zacatecas: 17 km S Villanueva (AVT), Laguna Balderama (= Valderrama) (Gidaspow, 1959: 283).

Notes: Brachypterous. It has been found in decidous forests and pastures, walking in the grassland or under stones, between 1100 and 2400m. Active adults have been captured from June to November (Erwin, 2007: 88)

Calosoma (Callitropa) atrovirens explanaticolle
Bates, 1891
Mexique, Oaxaca (type of Calosoma obscurum Géhin, 1885)
(coll. Muséum National d'Histoire Naturelle, Paris)
Calosoma (Callitropa) atrovirens explanaticolle
Bates, 1891
Mexico, Michoacan, env. Moralia, VII.2008, Lassalle leg.
Calosoma (Callitropa) atrovirens explanaticolle
Bates, 1891
Mexico, Michoacan, env. Moralia, VII.2008, Lassalle leg.


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