Calosoma (Callitropa) haydeni Horn, 1870

The systematic position of this species is still debated. Jeannel had in fact included it in Camedula, because its margin of the elytra is with evident serration. However, some other characteristics of external morphology, specially the squat and convex body shape, approach it to Blaptosoma, in which subgenus it has been included by Gidaspow.
haydeni is quite common in northern Mexico and can be found, although rare, in Texas, New Mexico and Arizona.
All authors distinguish two populations: haydeni haydeni and haydeni punctulicolle slightly different and geographically separated. However it is possible that some populations with intermediate characters exist in the contact area. Gidaspow (1959: 281) points out that she saw individuals related to one or the other of the two subspecies that were found in Texas, in two nearby locations, and Erwin (2007: 96) mentions C. haydeni haydeni from Chihuahua Mexico where does exist also C. haydeni punctulicolle.


Calosoma (Callitropa) haydeni haydeni Horn, 1870

Calosoma haydeni Horn, 1870: 69 (described from Colorado; syntype in Museum of Comparative Zoology, Harvard University, Cambridge, Ma)
Calosoma (Blaptosoma) laeve haydeni Breuning, 1928b: 45
Blaptosoma (s. str.) haydeni Lapouge, 1932: 392
Camedula (s. str.) haydeni haydeni Jeannel, 1940: 206
Calosoma (Blaptosoma) haydeni haydeni Gidaspow, 1959: 280

Length 21-23 mm. The northern populations are on average slightly smaller, with the sides of pronotum more rounded and narrower behind; moreover the striae on the elytra are more evident.
haydeni has been described from Colorado and it is also present in Arizona, Nevada, New Mexico and Texas. Erwin (2007: 96) quotes it from Chihuahua Mexico from where punctulicolle was described. Thus the hypothesis already formulated by Gidaspow of a partial mixing of the two populations in the contact area, would seem to be confirmed.

Examined specimens and literature’s data
Mexico. Chihuahua (Erwin, 2007: 96)
United States. Arizona: Mohave County (Gidaspow, 1959: 281); Colorado (type MCZ) Durango, San Juan mountains (Breuning, 1928b: 49; Nevada: Dorsey reservoir (Burgess & Collins,1917: 94); New Mexico: Torrance County (AVT), Koehler, Maxwell (Burgess & Collins,1917: 94); Texas: Brewster County, Alpine (Gidaspow, 1959: 281)

Notes: Brachypterous. It lives on field and pastures in mountains around an altitude of 2300m. Nocturnal, during the daytime seeks shelter under stones and debris. It feeds on caterpillars and pupae of lepidoptera. Active adults have been noted from April to November, and it is possible to find specimens overwintering in small cavities in the ground (Larochelle & Larivière, 2003: 178).
A brief description of larval stages can be found in Burgess & Collins (1917: 96).
The name of the species was proposed in honour of Ferdinand Vandeveer Hayden (1829 –1887) an American geologist, explorer and physician, famous for his pioneering surveying expeditions of the Rocky Mountains and later for his role in the establishment of Yellowstone National Park in 1872.

http://insects.oeb.harvard.edu/MCZ/
Calosoma (Callitropa) haydeni haydeni
Horn, 1870
United States, Col(orado) (Typus)
(coll. and photo: Museum of Comparative Zoology,
Harvard University, Cambridge, Massachusetts, USA)
http://insects.oeb.harvard.edu/MCZ/
Calosoma (Callitropa) haydeni haydeni
Horn, 1870
United States, New Mexico, Torrance county,
7ml E Willard, Laguna del Perro, 24.VII.1991,
Cassola leg. (coll. Vigna Taglianti)

Calosoma (Callitropa) haydeni punctulicolle Bates, 1891

Calosoma laeve punctulicolle Bates, 1891: 225 (type: Chihuahua, Santa Clara; British Museum of Natural History, London)
Calosoma (Blaptosoma) laeve punctulicolle Breuning, 1928b: 45
Blaptosoma (s. str.) laeve punctulicolle Lapouge, 1932: 392
Camedula (s. str.) haydeni punctulicolle Jeannel, 1940: 206
Calosoma (Blaptosoma) haydeni puncticolle Gidaspow, 1959: 281

Length 22-28 mm. The southern populations are slightly larger, and generally have less curved sides of pronotum while the intervals at the base of elytra are finely wrinkled but not scaly, as have the northern ones.
It is not rare to find in northern Mexico (Chihuahua, Coahuila, Durango, Monterrey, Nuevo Leon, Oaxaca, San Luis Potosi), moreover Gidaspow (1959: 281) saw one individual from Texas that she included in C. haydeni puncticolle, supposing that the two populations intergraded in the contact area.

Examined specimens and literature’s data
Mexico. Chihuahua: Agua Caliente, San Isidro, Catarinas, Jimenez, Santa Barbara (Gidaspow, 1959: 281), Santa Clara (Breuning, 1928b: 48), 4 km E of Pedernales, 37 km N of Temosachic, Minaca, Zaragoza, Cerro Venado, Ejido Zaragoza (UASM); Coahuila: La Gloria (Monclova) (Gidaspow, 1959: 281); Durango: Durango (MCZR), Durango city, Canelas, Promontorio (Breuning, 1928b: 48), Cuencame, Villa Ocampo, Santa María del Oro (Gidaspow, 1959: 281), El Pino, Nombre de Dios, Hidalgo del Parral (UASM); Monterrey (Breuning, 1928b: 48); Nuevo Leon: Puerto de Cineguillas, 2500 m (EM, SB); Oaxaca: Valle Nacional (UASM); San Luis Potosi: San Bartolo (www.gbif.org).
United States. Texas: Brewster County, Marathon (Gidaspow, 1959: 281)

Notes: Brachypterous. It lives on field and pastures from midlands up to an altitude of 2500m. Nocturnal, during the daytime seeks shelter under stones and debris. It feeds on caterpillars and pupae of lepidoptera. Active adults have been noted from May to November.

Calosoma (Callitropa) haydeni punctulicolle
Bates, 1891
Mexico, Nuevo Leon: loc. Puerto de Cineguillas, m 2500,
16.VII.1987, Migliaccio leg. (coll. Migliaccio)
Calosoma (Callitropa) haydeni punctulicolle
Bates, 1891
Mexico, Nuevo Leon: loc. Puerto de Cineguillas,
m 2500, 16.VII.1987, S. Bruschi leg.

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