Calosoma (Calodrepa) scrutator (Fabricius, 1775)

Carabus scrutator Fabricius, 1775: 239 (described from: Virginia); syntypes in Natural History Museum, London (Erwin, 1991: 23) and in Natural History Museum of Denmark, Copenhagen (Zimsen, 1964: 60)
Calosoma scrutator Fabricius, 1801: 213
Calosoma scrutator Breuning, 1927: 163
Calosoma (Calodrepa) scrutatrix Lapouge, 1932: 405
Calodrepa scrutator Jeannel, 1940: 77
Calosoma (Calodrepa) scrutator Gidaspow, 1959: 249
Calosoma (Calosoma) scrutator Erwin, 2007: 104


Length 25-35 mm. Among the species of the subgenre Calodrepa , C. scrutator is the one that, in some individual, reaches the largest adult size, although the size is very variable, depending also on the diet of the larvae.
As happens with C. aurocinctum, the elytra of C. scrutator are green with bronze luster and red lateral margin. However C. scrutator easily stands out because of its pronotum that is less narrowed behind, with surface roughness at its base, of dark blue or violet color, its sides with broad purple or golden green margin.
C. scrutator ranges across the entire of the United States but is more common in the eastern States. It has been found in southern Canada, still limited to the eastern part, and probably its reported presence in British Colombia must be considered exceptional. Towards south C. scrutator is present in Mexico and sporadically in some countries of Central America.
Gidaspow (1963: 283) quotes captures in Guatemala and at least one specimens has been recently found in El Salvador. It is present as small populations in Honduras, and possibly as an occasional visitor in Nicaragua. This would be confirmed by the attribution to C. scrutator (Van Den Berghe, 2016: 11) of the remains of a specimen found in the north of the country in the faeces of some predator and considered by the discoverers as remains of C. aurocinctum (Muñoz y Maes 2015), moreover one more specimens has been found not far away, near the border with Honduras. Finally, C. scrutator would also have been found on the coast of Venezuela (Gidaspow, 1963: 283) but in this case it is unlikely that it can be a permanent resident.

Examined specimens and literature’s data
Canada. British Columbia: Richmond (www.inaturalist.org/obs./25922777); New Brunswick (Bousquet, 2012: 231); Newfoundland (Breuning, 1927: 164); Nova Scotia: Sable Island (Bousquet, 2012: 231): Ontario: Middlesex, Ailsa Craig, (http://bugguide.net/), Prince Edward County, Grand Bend, Point Pelee National Park (Lindroth, 1961: 46), Dunks Bay, Toronto, Leamington (www.gbif.org/), Vaughan, (www.gbif.org), Otonabee-South Monaghan, St. Catharines, Murphys Point (www.inaturalist.org/); Quebec: Montreal (Breuning, 1927: 164)
El Salvador. La Libertad (www.inaturalist.org/obs./18084988)
Guatemala. Amatitlan, Huehuetenango (Gidaspow, 1963: 283)
Honduras. Francisco Morazán: Cerro Uyuca 1650m. (Van Den Berghe, 2016: 11, www.inaturalist.org/obs./ 15381341), Cerro Masicarán 800m. (Van Den Berghe, 2016: 11)
Mexico. Baja California (Leng, 1915: 565); Chihuahua: Pedernales, Ejido Zaragoza (UASM) ; Chiapas: Villaflores (www.gbif.org), Villa Corzo (inaturalist.org/observations/7347378); Guerrero: Chilapa (SB); Mexico DF.: Amanalco (www.inaturalist.org/observations/7604626); Michoacán: Morelia (https://www.inaturalist.org/obs.28597149); Morelos: Cirenio Longares (www.gbif.org); Nayarit: Tepic (UASM); Oaxaca: Juchatengo (UASM), Salina Cruz (Breuning, 1927: 164), Sierra de Ixtlan 2300 (http://www.ebay /), Santa Cruz Xoxocotlán (www.inaturalist.org/); San Luis Potosí (www.gbif.org); Sonora: Yecora (SB), Cibuta, Nacozari, Nácori Chico, (www.gbif.org), Aconchi, Moctezuma, San Nicolas, Mazatan, Cananea (UASM), Alamos, Magdalena de Kino (http://madrean.org/); Vera Cruz: Tuxpan, Playa Vicente (Breuning, 1927: 164)
Nicaragua. Esteli dep. Miraflor, loc. El Cebollal (sub aurocinctum Muñoz y Maes 2015: 5); Jalapa: Cerro Jesus, 1200m. (www.inaturalist.org/obs./ 6509785)
United States. Alabama: Tuscaloosa county (UASM), Montgomery County, Morgan County (bugguide.net/), Baldwin County (www.inaturalist.org); Arizona: Franklin County, Cienega creek (www.inaturalist.org), Santa Cruz County, Pima County, Cochise County (http://bugguide.net/), Maricopa County (http://madrean.org/); Arkansas: Bluff City (SB), Fayetteville (www.inaturalist.org/); California: Tulare co. (Burgess & Collins, 1917: 26), San Diego (www.inaturalist.org/); Colorado (Burgess & Collins, 1917: 26); Connecticut: Milford (www.inaturalist.org/); Delaware: Sussex County (http://bugguide.net/); District of Columbia: Boone County (http://bugguide.net/), Washington (Burgess & Collins, 1917: 27; www.inaturalist.org/); Florida: Tampa County, Polk County (http://bugguide.net/), Myakka River State Park (UASM), Orlando, Navarre, Pensacola, Tallahassee (https://www.inaturalist.org/); Georgia: Pickens County, Dawson County, Whitfield County (http://bugguide.net/); Illinois: Urbana (Burgess & Collins, 1917: 28), Chicago (SB), Riverside (SB), Hardin County, Cook County, (http://bugguide.net/), Jackson county (UASM); Indiana: Indianapolis (http://bugguide.net/), Terre Haute (www.inaturalist.org/); Iowa: Johnson County (http://bugguide.net/); Kansas: Sedgwick County (http://ninnescahlife.wichita.edu/), Shawnee County (http://bugguide.net/), Sumner County (http://madrean.org/); Kentucky: Letcher County, Pulaski County (http://bugguide.net/) Bowling Green, Fort Campbell (www.inaturalist.org/); Lousiana: Pitkin, Vernon Parish, Deridder (www.inaturalist.org/ ); Maryland: Glenburnie (EM), Howard County, Silver Spring (www.inaturalist.org/); Massachusetts: Dover, Cohasset, Onset (Burgess & Collins, 1917: 27), Brewster, Edgartown, Concord, Mackinac Island, Liberty (www.inaturalist.org/ ); Michigan (Burgess & Collins, 1917: 26); Mississipi: Oxford (www.jp-lawrence.com/); Minnesota: Ramsey co., Huston co., Fillmore co., Hennepin co. (Minneapolis), Lake Minnetonka (Gandhi & al,, 2005: 922); Missouri: Franklin County, Carter County (http://bugguide.net/), Vernon county (UASM), Peculiar (www.inaturalist.org/); Nebraska: Chase County, Lancaster County (http://bugguide.net/); Nevada: Vernon county (UASM); New Jersey (Burgess & Collins, 1917: 26); New Mexico: Water Canyon (Snow, 1881: 39), Santa Fe County (http://bugguide.net/), Albuquerque (www.inaturalist.org/); New York: Suffolk county (UASM); North Carolina: Buncombe County, Orange County (http://bugguide.net/), Bald Head Island, Raleigh, Asheville (www.inaturalist.org/); Ohio (Burgess & Collins, 1917: 26); Oklahoma: Stillwater (Burgess & Collins, 1917: 27), Okmulgee Co. (http://bugguide.net/), Mayes County, Tulsa, Coweta, Norman, Sawyer (www.inaturalist.org/); Pennsylvania: Philadelphia County (http://bugguide.net/), Pittsburgh, Bedford Co., Doylestown (www.inaturalist.org/); Rhode Island (Burgess & Collins, 1917: 26), Narragansett (www.inaturalist.org/ ); South Carolina: Laurens County (www.inaturalist.org/obs/28390151); Tennessee: Chattanooga (VV), Franklin County (http://bugguide.net/), Manchester (www.inaturalist.org/ ); Texas: Brazos county (SB), Smith county (SB), Willacy county (EM, SB), Starr county (EM), Nacogdoches county (SB), Curleson county, (SB), Tomball (SB), Fannin County, Bexar County, Harris County, (http://bugguide.net/), Kleberg county, Kerr county (UASM), Wise Co. Paradise (www.inaturalist.org) , Travis County (http://madrean.org/), Cooke Co. Valley View (www.inaturalist.org/obs/33199242), San Marcos, Lorena, Leakey, McKinney, Longview (www.inaturalist.org/); Utah: Washington County (www.inaturalist.org/obs/9102753); Virginia: Williamsburg, Albemarle County, (http://bugguide.net/), Belle Haven, Westmoreland County (www.inaturalist.org/obs).
Venezuela: Cumanà (Gidaspow, 1963: 283).

Notes: Winged, sometime diurnal but mostly nocturnal, attracted to light at night. Adults and larvae live mostly in forest and were reported feeding on various caterpillar as Alsophila pometaria, Paleacrita vernata, Malacosoma neustria, Erannis tiliaria, Alabama argillacea, Archips fervidana (Burgess & Collins 1917: 26). Adults have been observed in activity in the warm wet season from March to August with a maximum in April May. Depending on local climate condition, in the southern areas C. scrutator can remain active all the year long, or it has been found overwintering in the northern ones.
The larva was first described and figured by Chapuis & Candeze (1853: 371).

Calosoma (Calodrepa) scrutator
(Fabricius, 1775)
United States: Texas, Brazos county, College Station, april 1978
Calosoma (Calodrepa) scrutator
(Fabricius, 1775)
United States: Texas, Brazos county, College Station, april 1978

updated Jan. 12 2020

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