Calosoma (Calodrepa) scrutator (Fabricius, 1775)

Carabus scrutator Fabricius, 1775: 239 (syntype: Virginia; Natural History Museum of Denmark, Copenhagen)
Calosoma scrutator Fabricius, 1801: 213
Calosoma scrutator Breuning, 1927: 163
Calosoma (Calodrepa) scrutatrix Lapouge, 1932: 405
Calodrepa scrutator Jeannel, 1940: 77
Calosoma (Calodrepa) scrutator Gidaspow, 1959: 249
Calosoma (Calosoma) scrutator Erwin, 2007: 104


Length 25-35 mm. Among the species of the subgenre Calodrepa , C. scrutator is the one that, in some individual, reaches the largest adult size, although the size is very variable, depending also on the diet of the larvae. As happens with C. aurocinctum, the elytra are green with bronze luster and red lateral margin. However C. scrutator can be distinguished by its pronotum that is less narrowed behind, with surface roughness at its base, of dark blue or violet color, its sides with broad purple or golden green margin.
C. scrutator ranges across the entire of the United States but is more common in the eastern part. Moreover it occurs in southern Canada, and in Mexico, and towards south it could be sporadically present in some countries of Central America. Gidaspow (1963: 283) quotes captures in Guatemala and Van Den Berghe (2016: 11) assure that it is present as small populations in Honduras, and possibly as an occasional visitor in Nicaragua. In fact he believes that the remains of a specimen recently found there in the faeces of some predator and considered by the discoverers as remains of C. aurocinctum (Muñoz y Maes 2015), should be attributed to C. scrutator. Finally, C. scrutator would also have been found on the coast of Venezuela (Gidaspow, 1963: 283) but in this case it is unlikely that it can be a permanent resident.

Examined specimens and literature’s data
Canada: Ontario: Middlesex, Ailsa Craig, (http://bugguide.net/); Newfoundland; Quebec: Montreal (Breuning, 1927: 164); New Brunswick; Nova Scotia: Sable Island (Bousquet, 2012: 231)
Guatemala: Amatitlan, Huehuetenango (Gidaspow, 1963: 283)
Honduras. Francisco Morazán dep: Cerro Uyuca 1650m., Cerro Masicarán 800m. (Van Den Berghe, 2016: 11)
Mexico. Chihuahua: Pedernales, Ejido Zaragoza (UASM); Guerrero: Chilapa (SB); Nayarit: Tepic (UASM); Oaxaca: Salina Cruz (Breuning, 1927: 164), Sierra de Ixtlan 2300 (http://www.ebay/); Sonora: Yecora (SB), Nacozari, Aconchi, Moctezuma, San Nicolas, Mazatan, Cananea (UASM); Vera Cruz: Tuxpan, Playa Vicente (Breuning, 1927: 164)
Nicaragua. Esteli dep: Miraflor, loc. El Cebollal ( sub C. aurocinctum, Muñoz y Maes 2015: 5)
United States. Alabama: Montgomery County, Morgan County (http://bugguide.net/) Arizona: Cochise County, Santa Cruz County, Pima County, Cochise County (http://bugguide.net/); Arkansas: Bluff City (SB); Columbia: Boone County (http://bugguide.net/); Delaware: Sussex County (http://bugguide.net/); Florida: Tampa County, Polk County (http://bugguide.net/); Georgia: Pickens County, Dawson County, Whitfield County (http://bugguide.net/); Illinois: Chicago (SB), Riverside (SB), Hardin County, Cook County, (http://bugguide.net/); Iowa: Johnson County (http://bugguide.net/); Kansas, Shawnee County (http://bugguide.net/); Kentucky, Letcher County, Pulaski County (http://bugguide.net/); Maryland: Glenburnie (EM); Missouri, Franklin County, Carter County (http://bugguide.net/); Nebraska, Chase County, Lancaster County (http://bugguide.net/); New Mexico: Water Canyon (Snow, 1881: 39), Santa Fe County (http://bugguide.net/); North Carolina, Buncombe County, Orange County (http://bugguide.net/); Oklahoma, Okmulgee County (http://bugguide.net/); Pennsylvania, Philadelphia County (http://bugguide.net/); Tennessee, Chattanooga (VV), Franklin County (http://bugguide.net/); Texas: Brazos county (SB), Smith county (SB), Willacy county (EM, SB), Starr county (EM), Nacogdoches county (SB), Curleson county, (SB) Tomball (SB), Fannin County, Bexar County, Harris County, (http://bugguide.net/); Virginia: Williamsburg, Albemarle County (http://bugguide.net/)
Venezuela: Cumanà (Gidaspow, 1963: 283).

Notes: Winged, sometime diurnal but mostly nocturnal, attracted to light at night. Adults and larvae live mostly in forest and were reported feeding on various caterpillar as Alsophila pometaria, Paleacrita vernata, Malacosoma neustria, Erannis tiliaria, Alabama argillacea, Archips fervidana (Burgess & Collins 1917: 26). Adults have been found in the warm wet seasons from April to August and some specimens were found overwintering
The larva was first described and figured by Chapuis & Candeze (1853: 371).

Calosoma (Calodrepa) scrutator
(Fabricius, 1775)
United States: Texas, Brazos county, College Station, april 1978
Calosoma (Calodrepa) scrutator
(Fabricius, 1775)
United States: Texas, Brazos county, College Station, april 1978

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