Calosoma (Calodrepa) scrutator (Fabricius, 1775)

Carabus scrutator Fabricius, 1775: 239 (syntype: Virginia; Natural History Museum of Denmark, Copenhagen)
Calosoma scrutator Fabricius, 1801: 213
Calosoma scrutator Breuning, 1927: 163
Calosoma (Calodrepa) scrutatrix Lapouge, 1932: 405
Calodrepa scrutator Jeannel, 1940: 77
Calosoma (Calodrepa) scrutator Gidaspow, 1959: 249
Calosoma (Calosoma) scrutator Erwin, 2007: 104


Length 25-35 mm. Among the species of the subgenre Calodrepa , C. scrutator is the one that, in some individual, reaches the largest adult size, although the size is very variable, depending also on the diet of the larvae. As happens with C. aurocinctum, the elytra are green with bronze luster and red lateral margin. However C. scrutator can be distinguished by its pronotum that is less narrowed behind, with surface roughness at its base, of dark blue or violet color, its sides with broad purple or golden green margin.
C. scrutator ranges across the entire of the United States but is more common in the eastern part. Moreover it occurs in southern Canada, and in Mexico, and towards south it could be sporadically present in some countries of Central America.
Gidaspow (1963: 283) quotes captures in Guatemala and Van Den Berghe (2016: 11) states that it is present as small populations in Honduras, and possibly as an occasional visitor in Nicaragua. This would be confirmed by the attribution to C. scrutator of the remains of a specimen recently found in the north of the country in the faeces of some predator and considered by the discoverers as remains of C. aurocinctum (Muñoz y Maes 2015), moreover one more specimens has been recently found not far away, near the border with Honduras. Finally, C. scrutator would also have been found on the coast of Venezuela (Gidaspow, 1963: 283) but in this case it is unlikely that it can be a permanent resident.

Examined specimens and literature’s data
Canada. New Brunswick (Bousquet, 2012: 231); Newfoundland (Breuning, 1927: 164); Nova Scotia: Sable Island (Bousquet, 2012: 231): Ontario: Middlesex, Ailsa Craig, (http://bugguide.net/), Prince Edward County, Grand Bend, Point Pelee National Park (Lindroth, 1961: 46), Dunks Bay, Murphys Point , Toronto, Leamington (www.gbif.org/), Vaughan, (www.gbif.org); Quebec: Montreal (Breuning, 1927: 164)
Guatemala: Amatitlan, Huehuetenango (Gidaspow, 1963: 283)
Honduras. Francisco Morazán dep: Cerro Uyuca 1650m., Cerro Masicarán 800m. (Van Den Berghe, 2016: 11)
Mexico. Chihuahua: Pedernales, Ejido Zaragoza (UASM); Guerrero: Chilapa (SB); Nayarit: Tepic (UASM); Oaxaca: Salina Cruz (Breuning, 1927: 164), Sierra de Ixtlan 2300 (http://www.ebay/); Sonora: Yecora (SB), Nacozari, Aconchi, Moctezuma, San Nicolas, Mazatan, Cananea (UASM); Vera Cruz: Tuxpan, Playa Vicente (Breuning, 1927: 164)
Nicaragua. Esteli dep. Miraflor, loc. El Cebollal (sub aurocinctum Muñoz y Maes 2015: 5); Jalapa: Cerro Jesus (https://www.inaturalist.org/photos/8242124)
United States. Alabama: Tuscaloosa county (UASM), Montgomery County, Morgan County (bugguide.net/); Arizona: Cienega creek (www.inaturalist.org/observations/2837852), Santa Cruz County, Pima County, Cochise County (http://bugguide.net/), Maricopa County (http://madrean.org/); Arkansas: Bluff City (SB); California: Tulare co. (Burgess & Collins, 1917: 26); Colorado (Burgess & Collins, 1917: 26); Connecticut (Burgess & Collins, 1917: 26); Delaware: Sussex County (http://bugguide.net/); District of Columbia: Boone County (http://bugguide.net/), Washington (Burgess & Collins, 1917: 27); Florida: Tampa County, Polk County (http://bugguide.net/), Myakka River State Park (UASM); Georgia: Pickens County, Dawson County, Whitfield County (http://bugguide.net/); Illinois: Urbana (Burgess & Collins, 1917: 28), Chicago (SB), Riverside (SB), Hardin County, Cook County, (http://bugguide.net/), Jackson county (UASM); Iowa: Johnson County (http://bugguide.net/); Kansas: Sedgwick County (http://ninnescahlife.wichita.edu/), Shawnee County (http://bugguide.net/), Sumner County (http://madrean.org/); Kentucky: Letcher County, Pulaski County (http://bugguide.net/); Maryland: Glenburnie (EM); Massachusetts: Dover, Cohasset, Onset (Burgess & Collins, 1917: 27); Michigan (Burgess & Collins, 1917: 26); Mississipi (carabidae.org); Minnesota: Ramsey co., Huston co., Fillmore co., Hennepin co. (Minneapolis), Lake Minnetonka (Gandhi & al,, 2005: 922); Missouri: Franklin County, Carter County (http://bugguide.net/), Vernon county (UASM); Nebraska: Chase County, Lancaster County (http://bugguide.net/); Nevada: Vernon county (UASM); New Jersey (Burgess & Collins, 1917: 26); New Mexico: Water Canyon (Snow, 1881: 39), Santa Fe County (http://bugguide.net/); New York: Suffolk county (UASM); North Carolina: Buncombe County, Orange County (http://bugguide.net/); Ohio (Burgess & Collins, 1917: 26); Oklahoma: Stillwater (Burgess & Collins, 1917: 27), Okmulgee County (http://bugguide.net/); Pennsylvania: Philadelphia County (http://bugguide.net/); Rhode Island (Burgess & Collins, 1917: 26); Tennessee: Chattanooga (VV), Franklin County (http://bugguide.net/); Texas: Brazos county (SB), Smith county (SB), Willacy county (EM, SB), Starr county (EM), Nacogdoches county (SB), Curleson county, (SB) Tomball (SB), Fannin County, Bexar County, Harris County, (http://bugguide.net/), Kleberg county, Kerr county (UASM), Travis County (http://madrean.org/); Utah: Washington County (www.inaturalist.org/observations/9102753); Virginia: Williamsburg, Albemarle County, (http://bugguide.net/).
Venezuela: Cumanà (Gidaspow, 1963: 283).

Notes: Winged, sometime diurnal but mostly nocturnal, attracted to light at night. Adults and larvae live mostly in forest and were reported feeding on various caterpillar as Alsophila pometaria, Paleacrita vernata, Malacosoma neustria, Erannis tiliaria, Alabama argillacea, Archips fervidana (Burgess & Collins 1917: 26). Adults have been found in the warm wet seasons from April to August and some specimens were found overwintering
The larva was first described and figured by Chapuis & Candeze (1853: 371).

Calosoma (Calodrepa) scrutator
(Fabricius, 1775)
United States: Texas, Brazos county, College Station, april 1978
Calosoma (Calodrepa) scrutator
(Fabricius, 1775)
United States: Texas, Brazos county, College Station, april 1978

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