Calosoma (Calodrepa) wilcoxi LeConte, 1848

Length 17-22 mm. C. wilcoxi has, on average, smaller dimensions when compared to all the other species collected into Calodrepa. At first glance it looks a small C. scrutator, because of the body shape and color, but it is easily recognized because it is the only species of the group that has the metatrochanters bearing a seta and that has straight or, at most, slightly arcuate, male's mesotibiae.
A few aberrant specimens of C. wilcoxi, having dark brown or decidedly dark blue-black upperbody instead of the typical metallic green, are found occasionally in Texas. Two of these aberrant specimens were published by Ray & al. (2016).
C. wilcoxi ranges across most of the eastern United States from the Atlantic coast to the Great Plains. It is found in Ontario and Quebec (in this last case probably only it as an occasional visitor) down to Texas, but it has never been recorded from Mexico, and its presence in California indicated by Burgess & Collins (1917: 39), is considered unlikely by Bosquet, (2012: 230).
C. wilcoxi was imported into Hawaii, as a potential agents of biological control with some more American species, but without success (Liebherr et al. 2023).

Examined specimens and literature’s data
Canada. Ontario: Toronto (www.inaturalist.org/obs/12972148), Prince Edward County, Ottawa, Point Pelee National Park (Lindroth, 1961: 47), Leamington (BIOUG), Rouge Park (UASM), Murphys Point (www.gbif.org), Norfolk County (www.inaturalist.org); Quebec: Aylmer (Lindroth, 1961: 47).
United States. Alabama: Boaz (www.gbif.org/), Madison county, Tuscalosa county, Marshall county (www.inaturalist.org); Arkansas: Fort Smith, Jonesboro, Holland (www.inaturalist.org); California: Tulare co. (Burgess & Collins, 1917: 39); Connecticut: Hamden (www.inaturalist.org); Delaware: Dewey Beach (www.inaturalist.org); District of Columbia: Washington (www.inaturalist.org); Georgia: Floyd co., Dade co. (www.inaturalist.org); Illinois: Peoria county, Peoria (EM, SB), McDonough county, Macomb (UASM), Jackson county, Carbondale (OSUC), Champaign County (http://bugguide.net/); Indiana: Evansville, Bloomington (www.inaturalist.org); Iowa (Burgess & Collins, 1917: 38): Fremont Co. (www.inaturalist.org); Kansas: Crawford, 3 mi NE Pittsburg (SEMC), Johnson County (http://bugguide.net/); Kentucky: Rockcastle Co. (OSUC), Christian county (UASM); Louisiana: Saint John the Baptist Parish, Saint Tammany Parish (Bousquet, 2012: 232); Maine (Bousquet, 2012: 232); Maryland: Allegany county, 2ml W Pawlings (EM), Mongomery co. (https://bugguide.net/), Dorchester Co., Baltimore City (www.marylandbiodiversity.com), Cove Point (http://www.covepoint-trust.org/), Garrett County, Frederick county (www.inaturalist.org); Massachussets: Fairhaven (www.inaturalist.org); Michigan: Washtenaw Co., Ann Arbor (OSUC); Mississipi (Bousquet, 2012: 232): Lafayette County, Oktibbeha County (www.inaturalist.org); Missouri: Columbia (EM), Taney Co., Branson (OSUC), Adair County (www.inaturalist.org); Minnesota (Bousquet, 2012: 232); Nebraska: Namaha county, Brownville (SB), Sarpy county, Bellevue (SB), Lincoln (EM); New Hampshire: Rockingham county (Bousquet, 2012: 232), Cheshire County (www.inaturalist.org); New Jersey: Atlantic county, (OSUC), Cape May county (www.inaturalist.org); New York: Ithaca (www.inaturalist.org), Suffolk county, East Hampton, Long Island (EM), (OSUC), Queens County (http://bugguide.net/); New Mexico: Cibola County (http://bugguide.net/); North Carolina: Belmont, Jordan Lake (www.gbif.org/), Mecklenburg county, Johnston county, Union county (www.inaturalist.org); North Dakota (Bosquet, 2012: 232); Ohio: Lake Co., Mentor (OSUC), Hamilton Co. (OSUC), Adams Co. (OSUC), Wayne Co. (OSUC), Clifton (OSUC), Columbus (OSUC), Franklin Co., Blendon Twp. (OSUC), Delaware Co. (OSUC), Ashtabula Co., Ashtabula, (OSUC); Oklahoma: Pontotoc County (http://bugguide.net/), Mayes County, Lincoln County, Pawnee County (www.inaturalist.org); Pennsylvania: Washington County, Forest Hills, Gettysburg (www.inaturalist.org); Rhode Island (Burgess & Collins, 1917: 38); South Carolina: Richland county (www.inaturalist.org), South Dakota (Bosquet, 2012: 232); Tennessee: Lake county, Reelfoot Lake (UASM), Chester County, (http://bugguide.net/); Texas: Brazora county (Bousquet, 2012: 232); Brazos county, College Station, (OSUC, SB), Brown County, Travis County, (http://bugguide.net/), McLennan County (Ray & al., 2016), Coryell County, Tarrant County, Wise county (www.inaturalist.org); Utah: Gunnison Butte (www.gbif.org/); Virginia: Richmond City county, Chesterfield county, Campbell County (www.inaturalist.org); Washington, DCA (UASM); West Virginia, Morgan County (http://bugguide.net/); Wisconsin: Lafayette County (http://bugguide.net/), La Crosse, Racine, Richland, Winnebago (Messer, 2009: 33), Dane County (www.inaturalist.org).

Notes: Winged, it is attracted to light at night but it is diurnal as well. C. wilcoxi is a good climber and it is not rare to observe large gathering of adults feeding upon species of Geometrid caterpillar.
Most of the active individuals were captured from April to June, that is also the coupling period. Any way, according with local climate, C. wilcoxi seems to be active all year round, while in the northern areas, it may happen to find adults overwintering in small cavities in the ground.
There is no explicit dedication of this species and it is difficult to identify the person which the name “Wilcoxi” refers to. However, in other publications of the author, there are some references to a “Mr. Wilcox” who gave him various beetles of Illinois.
A first brief description of larval stages can be found in Burgess & Collins (1917: 41).

Calosoma (Calodrepa) wilcoxi ♂ LeConte, 1848 United States: Texas, Brazos county, College Station, 10 april 78

Calosoma (Calodrepa) wilcoxi ♀ LeConte, 1848 United States: Texas, Brazos county, College Station, 10 april 78

Calosoma (Calodrepa) wilcoxi ♂ LeConte, 1848 United States: Texas, Wise county, Annaville, July 13 2001 (image from: www.inaturalist.org/obs/41339622)

Calosoma (Calodrepa) wilcoxi ♀ LeConte, 1848 United States: Texas, Wise county, Boyd, June 8 2001 (image from: www.inaturalist.org/obs/10031851)

Calosoma (Calodrepa) wilcoxi ♂ LeConte, 1848 United States: Texas, McLennan County, Waco, Baylor University Campus, April 1, 2012 (coll. Matthias Seidel)

Calosoma (Calodrepa) wilcoxi ♀ LeConte, 1848 United States: Texas, McLennan County, Waco, Baylor University Campus, April 1, 2012 (coll. Matthias Seidel)