Calosoma (Calosoma) sycophanta (Linné, 1758)

Carabus sycophanta Linné, 1758: 414 (described from Europa) type material: unspecified number of specimens; possible syntypes in Museum of Evolution Zoology, Uppsala University (UUZM, Catalogue of Linnaean specimens 2001: 21) and Linnean Society of London, (Lindroth, 1957: 334)
Calosoma sycophanta Weber, 1801: 20
Calosoma sycophanta var. severum Chaudoir, 1850: 422; (described from Anatolie) holotype by monotypy ♂ designated by Deuve (1978: 248) in Muséum National d'Histoire Naturelle, Paris [examined]
Calosoma sycophanta var. nigrocyaneum Letzner, 1850: 95; (distribution: Schlesien) original material: unspecified number of specimens, original specimen(s) in Senckenberg Deutsches Entomologisches Institut, Müncheberg (Casale & al, 1982: 79)
Calosoma sycophanta var. marginatum Letzner, 1850: 95 (idem)
Calosoma sycophanta var. azureum Letzner, 1850: 95 (idem)
Calosoma sycophanta var. purpureoaureum Letzner, 1850: 95 (idem)
Calosoma sycophanta var. cupreum Letzner, 1850: 95 (idem)
Callipara sycophanta Motschoulsky, 1865: 308
Callipara sycophanta rapax Motschoulsky, 1865: 309 (described from environs de Amasia) type material: 1 ♂, repository not stated.
Calosoma sycophanta var. smaragdinum Rossi, 1882: 204; (distribution: Hungary) possibly some original specimen in Museum für Naturkunde, Berlin (Casale & al, 1982: 79)
Calosoma sycophanta var. Habelmanni Schilsky, 1888: 180 (described from Berlin)
Calosoma sycophanta var. purpuripennis Reitter, 1891: 257; (described from Margelan) type in Magyar Természettudományi Múzeum, Budapest (Jeannel, 1940: 82)
Calosoma sycophanta ab. corvinum Heller, 1897: 312
Calosoma sycophanta ab. anthracinum Houlbert, 1907: 292; (described from: La Baule) (preoccupied by anthracinum Dejean 1831)
Calosoma sycophanta ab. prasinum Lapouge, 1907: 124; (distribution: South Azerbaijan, North Iran, North Iraq) type material: 3 ♀ 1 ♂; lectotype ♀ (Elbourz, Talyche) designated by Deuve (1978: 255) in Muséum National d'Histoire Naturelle, Paris [examined]
Calosoma sycophanta ab. solinfectum Jänichen, 1914: 298 (described from Brandenburg, Bärwalde) holotype in Senckenberg Deutsches Entomologisches Institut, Müncheberg (Döbler, 1975: 141)
Calosoma sycophanta sycophanta Breuning, 1927: 173
Calosoma sycophanta var.? lapougei Breuning, 1927: 174 (new name for anthracinum Houlbert, 1907)
Calosoma sycophanta nigrosuturale Jacquet, 1930 (cited in Lorenz, 2005: 68)
Calosoma (Callipara) sycophanta, Lapouge, 1932: 403
Calosoma sycophanta nigroaeneum Polentz, 1937: 7
Calosoma sycophanta var. Hankói Narozsny, 1938: 75 (described from Gyulaj)
Calosoma sycophanta var. Gizellae Narozsny, 1938: 75 (described from Gyulaj)
Calosoma (Calosoma) sycophanta, Jeannel, 1940: 81
Calosoma (Calosoma) sycophanta subsp. prasinum, Jeannel, 1940: 82 (distribution: Talisch et Elbourz)
Calosoma (Calosoma) sycophanta subsp. purpureipennis, Jeannel, 1940: 82 (distribution: Turkestan)
Calosoma (Calosoma) sycophanta subsp. severum, Jeannel, 1940: 82 (distribution: Asie Mineure et Caucase)
Calosoma (Calosoma) sycophanta altaica Obydov, 2010: 62 (type locality: western Altai, Zmeinogorsk distr.) type material: 3 ♀♀, holotype in Museo di scienze Naturali "Cavalier Locca", Guardabosone

Length 21-35 mm. C. sycophanta is characterized by a strongly transverse pronotum, sinuate behind. The lateral margin of the pronotum is thick, conspicuously elevated, and it disappears just before of the basal angles. Head and pronotum are wrinkled and sparsely punctured. The sculpture of elytra is flattened with smooth intervals and with clearly visible punctures on the striae. The tarsi of the male forelegs have hairy pads under the sole of the first three dilated segments.
C. sycophanta is found, and in some years is quite common, in northern Africa and throughout Europe (including the Mediterranean Islands).
Towards north, C. sycophanta has been found sporadically in Denmark. In Sweden it is also very rare and it has been captured in single specimens only in the south of the county (Lindroth, 1985: 47), but some remains of this species were found in lacustrine sediment (gyttja) deposited on the shore of the ancient lake Bare, near Halmstad and dated by pollen analysis to a warmer period around 10,000 years BC (Welinder, 1970). Recently it has been accidentally imported in Finland, with vegetable foods (Bistrom, 1983: 212). In Norway, any recent captures of C. sycophanta are not known and in any way they seem improbable, however a single specimen found in 1921 should still be preserved in the entomological collections of the University of Bergen (
To the east, C. sycophanta spreads to Asia Minor, Iran, central Asia (Azerbaijan, Turkmenistan and Kyrgyzstan) up to western Siberia. It is indicated by various authors (Yu, 1982; Deuve, 1997) of Xinjiang in northern China, based on an ancient citation of Breuning, (1927: 174), which literally reads: “ Turkestan, towards East up to the Thian Shan”.
C. sycophanta is not rarely found on the south-eastern coast of England but it is possibly an occasional immigrant. It has been imported into the eastern United States in 1905-1910 (Burgess 1911, Burgess & Collins 1917). Its range has continued to expand at a rate of dispersion, calculated at 6 km/year and now extends from southern Maine and all New England states south into Pennsylvania, Delaware, Maryland, and West Virginia (Schaefer & al. 1999). C. sycophanta was unsuccessfully introduced in Canada, in New Brunswich and in Vancouver island (Lindroth, 1961: 46), however, it seems to be present in Nova Scotia (Erwin, 2007: 105).
Normally the upper body of C sycophanta is two-tone with bluish head and pronotum, and with coppery or golden green elytra. However the color is variable, in some cases also because affected by climatic conditions under which the pupation took effect. It is possible to find specimens with cupric elytra and dark bluish pronotum or, very rarely, it is possible to find uniformly colored specimens, green blue (prasinum Lapouge) or light blue (azureum Letzner). In some localities of southern Turkey, with relative frequency, are found specimens with a uniform purplish brown or black color (severum Chaudoir, rapax Motschoulsky), similar to the rare individual forms that occasionally have been encountered in various localities of central Europe and that were differently named (habelmanni Schilsky, corvinum Heller, anthracinum Houlbert, lapougei Breuning).
Breuning (1927: 171) assumed all these forms to be synonyms of the typical form as they cohabit with this. Yet, he considered the black form (anthracinum renamed lapougei) of Brittany worthy of name. On the other hand, Jeannel (1940: 81) re-evaluated the three chromatic forms: severum of southern Turkey, prasinum of southern Azerbaijan and northern Iran, and purpuripenne of Uzbekistan, as subspecies as they would have constant characteristics in these regions.
However we believe that C sycophanta is a species that can present individuals with variable color, sometimes more numerous or predominant in some geographical areas, but that are always in mixed populations and therefore that these populations do not deserve to be distinguished at the subspecific level.
The same applies to the only subspecies described in recent times (sycophanta altaica). According to its description (Obydov, 2010: 62), this new taxon should be distinguished by a set of very weak and uncertain characters such as less accentuated micro-sculpture of head and pronotum, less transverse pronotum and elitrae slightly shorter and broader, with some barely perceptible difference in the color of the upper body.

Examined specimens and literature’s data
Albania. Sauk, Voskopojë, Shkodër, Borshi near Vlorë (Gueorguiev, 2007: 30), Durrës (
Algeria. Le Telagh near Daya, forêt de Blidah, forêt de Teniet-el Had, forêt de Boghari, forêt de Akfadou (Kabilie), Sain- Charles, forêt de Batna, Bone, forêt de l’Edough, Bou-Mzeran (Bedel, 1895: 19)
Armenia. Kosrov reserve (VV)
Austria. Burgenland: Loretto am Leithagebirge, Neusiedl am See, Purbach am Neus (Mandl 1957: 106); Podersdorf, Nickelsdorf, Zitzmannsdorfer Wiesen (; Kärnten: Leopoldskirchen, Loibltal, Sagritz im oberen Mölltal (Mandl 1957: 106), Sagritz, Heiligenblut (Pacher, 1853); Niederösterreich: Leithagebirge, Mödling, Niederweiden, Oberwaltersdorf, Plank, Straßhof, Trumau bei Baden, Vöslau, Wiener Neustadt (Mandl 1957: 106), Perchtoldsdorf (; Steiermark: Admont, Graz (; Tirol: Brandenberg, Innsbruck-Umg., Judenstein im Ahrntal bei Innsbruck, Jenbach, Lechgebiet, Gramais, between Matrei and Pfons, Obernberg am Inn, Ötz (Mandl 1957: 106), Wiesing, Hötting b. Innsbruck (; Steiermark: Admont, Plabutsch bei Graz (Mandl 1957: 106), Pürgg (; Oberösterreich: Obernberg am Inn, Urfahr, Grünburg (; Vorarlberg: Bludenz; Bregenz; Feldkirch (Mandl 1957: 106); Wien: Wien-Lainzer Tiergarten, Perchtoldsdorf, Vorder Hainbach, Sofienalpe (Mandl 1957: 106)
Azerbaijan. Elbourz, (lectotype of Calosoma prasinum MNHN), Talysch (Breuning, 1927: 173), Azerbaijan (Ebay)
Belarus. Białowieża Forest (
Belgium. Gent, Chaudfontaine (
Bosnia-Erzegovina. Domanovići, Stolac, Volujak Mt. (Apfelbeck, 1904 : 16), Zavidovići (sub var. severum, Narozsny, 1938: 75), Ljubinje (
Bulgaria. Sliven, Stanimaka (Asenovgrad), Burgas (Apfelbeck, 1904 : 16); Krichimska Kuriya, Brodilovo Vill. (Buresch & Kantardjieva, 1928: 64); Varna (SB), Kalimantsi (EM), Belogratchik (; Kostenets, Sturgach Planina (Guéorguiev & Guéorguiev, 1995: 40); Bulgari Vill., Ahtopol, Bliznak Vill., Maslen nos Cape, Ropotamo, Strandzha, Gramatikovo Vill., Slantschew brjag (Burgas), Slivarovo Vll. (, Haskovo (Buresch & Kantardjieva, 1928: 64), Harmanli (Guéorguiev, 2004: 382), Bryanovishtitsa Hut 930m, Belovo Pamporovo, along Chepinska river and Alabak (Guéorguiev & Lobo, 2006: 288), Lozenets, Sozopol (Tsentar) (
Croatia. Izola, (Isola d’Istria), Labin (Albona), Rijeka (Fiume), Platak, Mt. Učka (Maggiore) Losinj (Lussino), Unije (Unie) (Müller, 1926: 44); Vrlika, Crikvenica, Pula (, Merag, Šibenik (
Cyprus. (Breuning, 1927: 174)
Czech Republic. Uničov Břevenek (VV), Moravia (sub azureum det. Mandl; NMB), Česká Lípa, Horní Dohalice, Neratovice, Kolín, Benešov u Prahy, Pátek, Moravský Krumlov, Pouzdřany, Valtice ( ; Břevenec (; Brno: Jihomoravský kraj (
Denmark. Jutland, Sønderjylland (Schiødte, 1841: 308); Skellebjerg (Lindroth, 1985: 47)
Estonia Viljandi (Haberman, 1968: 117)
Finland. Uusimaa: Kauniainen (Bistrom, 1983: 212)
France. Alpes-de-Haute-Provence: Digne-les-Bains (; Alpes-Maritimes: Saint-Étienn e-de-Tinée, Saint-Dalmas-le-Selvage (; Ardèche: Résurgence de Font vive; Bouches-du-Rhônes: Montagnette, Vallen Pont d'Arc (; Calvados: Forêt de Cerisy (; Corse: Corte (, Portovecchio (SB); Gard: Saint Jean du Gard, Dions; Haut-Rhin: Dessenheim, Rixheim; Hérault: Montarnaud (, Ganges (EM); Loire-Atlantique: La Baule, Le Pouliguen (var. lapougei Jeannel, 1940: 82); Lot: Cressensac (; Provence: Avignon (EM); Midi-Pyrénées: Pibrac (; Pyrénées-Atlantiques Sare (, Saint Jean de Luz, Biarritz, Eaux Bonnes, Buzy, Bois de Sauvagnon (Jeanne, 1984: 66); Pyrénées-Orientales: Òpol (; Rhône: Lyon (; Var: Port Grimaud (, Collobriéres (, Aiguines, Bormes-les-Mimosas (, Frejus (; Vaucluse: massif du Luberon (, Les Mayons (EM); Yvelines: Gommecourt (
Germany. Bavaria: Wasserlos (; Bayern: Königsdorf (; Brandenburg: südlich Brandenburg (sub azureum det. Mandl; NMB), Baruth (Weisskeissel), Woltersdorf, Niedergörsdorf (; Hesse: Eschwege, Schweinsberg, Mainhausen (; Niedersachsen: Harz (; North Rhine-Westphalia: Köln (; Rhineland-Palatinate: Taunus, Bleidenstadt, Nassau, Kirn, Frei-Laubersheim (; Saarland: Perl (; Schleswig-Holstein: Kiel (Schiødte, 1841: 308), Glinde (; Thüringen (Hartmann, 2007: 159)
Greece. Mt. Olympus (EM), Mt. Parnassós (Apfelbeck, 1904 : 16); Peloponnesus: Pylos-Nestoras (Koubeh Forest), Mt. Taygetos (Apfelbeck, 1904 : 16), Laconia: Kariovouni (, Ahaia Kalogria (EM), Kalavrita (EM), Nestos gorge (, Dhomokos (, Creta island: (Apfelbeck, 1904 : 16), Archanes-Asterousia, Lasithi ( 14712357); Thessaloniki: Ano Poli (
Hungary. Gyulaj, Miskolc, Diósgyőr, Lillafüred, Fehérgyarmat, Nyíregyháza, Debrecen, Pécel, Budapest-Pestszentimre, Budapest-Rákosszentmihály, Szokolya, Sopron, Kőszeg, Mariabesnyo, (Narozsny, 1938: 75)
Iran. Mazandaran prov: Marzanabad (EM, SB), Siah-bishe, (Azadbakhsh & al. 2015: 45); Gilan prov: Rasht distr (SB); Elburz, Mts Talych (Ardabil or Gilan provinces) (type of prasinum; MNHN); Kermanshah (Lourestan) prov: Kermanshah 1470m. (sub prasinum Lapouge, 1907: 124).; Golestan prov: Nokandeh (Ghahari & al., 2009: 439).
Israel. Haifa: Mont Hermon 1600 m (EO)
Italy. Abruzzo: Monte Fiore (Bruno, 1974), L’Aquila, Cerchio (Leoni, 1909: 133), Pescasseroli, Val Fondillo, Fallo, Prata d'Ansidonia, Raiano (AVT), Monti della Difesa, Monte Schiena Cavallo (Luigioni, 1933: 5); Basilicata: Fiume Basento, Maratea (CKmap), Viggiano (AVT), foresta di Gallipoli Cognato (; Calabria: Aspromonte Monte Pecoraro Monte Pollino (Magistretti, 1965: 59), Molochio, San Giorgio Morgeto (AVT); Campania: Napoli, Quarto Caselle (AVT); Emilia: Bologna, Imola, Ladino, Pineta di Ravenna, Cervia (Magistretti, 1965: 58), Casinalbo, Mesola (Magistretti, 1968:183), Lago Brasimone, Ladino, Bosco di Scardavilla, Meldola (AVT), Piacenza (AC), Baricella, Monte Gottero, Monte Maggio, Spilamberto (CKmap); Friuli: Tarvisio (SB), Udine, Tarcento, Grado, Monfalcone, Aidussina, Monrupino, Lipizza (Müller, 1926: 44), Lago di Doberdò (Magistretti, 1965: 58), Opicina, Basovizza (SB), Sistiana (Battiston & Biondi, 2015: 7), Aurisina, Slivia, Lago di Selva (AVT), Gropada (CKmap); Lazio: Roma (SB), Bagnoregio (SB), Parco Naz. Circeo (AVT, EM), Filettino (EM), Cecchina (SB), Manziana (SB), Monterotondo (EM), Mt. Tagliente (EM), Frascati (EM), Colonna (EM), Bolsena, Civitavecchia, Mt Autore, Marino, (Magistretti, 1965: 58); Acciarella, Torre Astura, Cerasella, Monti della Duchessa, Monte Gennaro, Ienne, Lavinio, Foce Verde, Monte San Biagio, Nemi, Percile, Monte Pizzuto, Rocca di Papa, Monte Antenne, Monte Rufeno, Monte Faito, Terracina (AVT), Acilia, Monte Fogliano, Lavinio, Mentana, Pomezia, Rocca Priora, Santa Severa, Sant'Angelo Romano (Bruno, 1974) Ronciglione, Ariccia, Anzio (Luigioni, 1899: 212); Liguria: Pontedecimo (GKmap), Conna (GA), Genova (Magistretti, 1965: 58), Altare, Passo Muratone (AVT); Lombardia: Valtellina (EM), Ponte San Pietro (Breuning, 1927: 173), Milano, Monza, Casatenovo, Maccagnoarcisate, Turbigo, Melegnano, Ottobiano, Spino d’Adda, Como, Bellagio, Dervio, Premana, Mt. Legnone, Bagni del Masino, Chiesa Valmalenco, Val di Arigna, Ponte in Valtellina, S. Caterina Valfurva (Magistretti, 1965: 58), Pozzolengo Cantalupo (Battiston & Biondi, 2015: 7) Arigna (AVT), Maccagno (GA), Arona (; Marche: Acquacanina Bolognola Cingoli Sassoferrato Visso (ex coll. Battoni) Sefro (, Fiordimonte Pievebovigliana (AVT), Carpegna, Fabriano Monte Palazzo Borghese (Magistretti, 1965: 58), Campodonico, Colle di Fossato Potenza Picena (Magistretti, 1968 183); Molise: Campitello Matese (EM); Piemonte: Viozene, Rivoli, Pinerola, Campiglione Fenile, Bollengo, Colle del Turlo (Magistretti, 1965: 58), Limone 2180m (www.floraitaliae), Viozene, Sambuco, Rifugio Dante Livio Bianco (AVT), Antronapiana, Bienca, Colle della Boaria, Burolo, Candia Canavese, Chiaverano. Domodossola, Ivrea, Netro, Pella, Rimasco, Rueglio. Torre Canavese (AC); Trino, Vico Canavese, Frassineto Po (GA); Alba Cissone,Dogliani Grignasco Monforte d'Alba Pagno Pocapaglia Revello (CKmap); Puglia: St. Basilio Mottola, Noci (Magistretti, 1965: 58), Castellaneta (AVT), Gioia del Colle (Marcuzzi, Turchetto Lafisca, 1977); Sardegna: Pula, Is Cannoneris (, Tempio Pausania, Orgosolo, Aritzo, Seui, Macomer, Torre di Flumentorgiu, Carloforte (Magistretti, 1965: 58), Iglesias, Mogorella, Nuoro (Magistretti, 1968: 183), Baunei, Cagliari, Cuglieri, Desulo, Isola Foradada, Isola La Maddalena, Isola Mal di Ventre, Isole de Li Nibani, Monte Is Pauceris, Mannu, Isola San Pietro, Isola Paduleddu, Santadi, Monte Sette Fratelli, Isola Soffi, Villanova Monteleone (AVT), Berchidda, Dorgali, Monte Limbara, Isolotto Camize, Monte Spada, (AC), Isola Asinara, Cala Gonone, Isola Molarotto, Isola Mortorio, Isola La Presa, Isola Piana di Alghero, Isola Il Toro (MGe), Isola Sant'Antioco (Piras, Pisano, 1972), Loiri Porto San Paolo (, Urzulei Cantoniera Bidicolai (coll. Zilli); Sicilia: Monte Pagano (, Castelbuono (Ragusa, 1883: 13), Guidomandri (Vitale, 1912: 201), Geraci Siculo (Magistretti, 1965: 58), Monte Pagano (; Sud Tirolo: Brunico (Magistretti, 1968: 183), Bolzano, Campodazzo, Bressanone, Naturno, San Felice, Vistrad, San Leonardo (Val Passiria), Stava, Val d'Ultimo, Val Venosta (Peez Von, Kahlen, 1977), Bo Fiè allo Sciliar (Voels am Schlern), Passo del Rombo (Timmelsjoch), Mt. Sonnenberg (; Toscana: Isola d'Elba, Marciana (, Marsiliana (AVT), Isola del Giglio (Gridelli, 1926: 433), Forte dei Marmi, Monte Amiata (Magistretti, 1965: 58) Siena (Magistretti, 1968:183), Firenze, Lucca, Bagni di Lucca, Montecchio, Mugello, Val di Nievole, Pisa, Querceto (Piccioli, 1869: 207), Sansepolcro (Gagliardi, 1941: 92), Vallombrosa (Cecconi, 1897); Trentino: Monte Baldo, Zambana, Val di Non (Magistretti, 1965: 58), Cavalese, Nogarè, Rovereto, Val di Sole, Trento, Torcegno (Bertolini, 1887: 88), Isera, Lizzana, Pozzacchio, St. Ilario, (Halbherr, 1885: 7), Levico Terme (CKmap); Umbria: Foligno, Nocera Umbra, Orvieto (Leoni, 1909), Norcia, Castelluccio, Mt Palazzo Borghese, Pistrino (Magistretti, 1965: 58), Terni, Icciano (AVT), Lippiano (Gagliardi, 1941); Valle D’Aosta: Valtournanche (Magistretti, 1965: 58), Colle del Gran San Bernardo (AC); Veneto: Verona (EM), Monte Baldo, Caprino Veronese, Padova (Magistretti, 1965: 58), Venezia Lido (CKmap), Colli Berici, Colli Euganei, Castelgomberto, Parco delle Grotte di Oliero, Cortina d'Ampezzo, Falcade (Battiston & Biondi, 2015: 7)
Kazakhstan. lake Zaysan reg. (Wernoje), Tien Shan mountains (Breuning, 1927: 173), Almaty prov: Aksu (SB), Enbekchikazakh, Sayram-Ugam N.P. (
Kyrgyzstan. Fergansky mts, Uruni-Bash 1600 m (SB); Fergana Mt. range, Isfan-Dzhailoo Mts. (http://www.ebay /); Boo-Tek river (
Latvia. (Telnov, 2004: 8)
Liechtenstein. (
Lithuania. (Tamutis & al., 2011)
Luxenbourg. Vianden, Wintrange (
Macedonia. (
Mauritania. Titen Yaya (EM)
Moldavia. (
Montenegro. Stanišići (, Komovi Mt. (Apfelbeck, 1904 : 16)
Morocco. Meknès-Tafilalet, Ifrane (EM); Eastern Region, Berkane (EM); Grand-Atlas Central: Talmest 2600m., Tassaout n'Ouidamen 2200m. (Kocher , 1938 : 78); Melilla; Muley Rechid; Kariet Arkmane; Jbel Kouali, Taourirt; (Chavanon & François 2014: 201);, beach of Tanger (Bedel, 1895: 19); Moyen Atlas (
Netherlands. Gelderland: Nijmegen; Friesland: Sneek; Zuid-Holland: Den Haag, Loosduinen, Noordwijk; Limburg: Venlo (NBC,
Norway. Nord-Trøndelag: Snåsa (Brønstad) (
Poland. Pomerania: Szklana Huta, Miastko (; Łódź: Rezerwat Spała, (; Silesia: Klobuck, Bytom (; Cracovia: Zwierzyniec (
Portugal. Bussaco, Coimbra, Serra de Estrella, Cintra (Oliveira, 1876: 21)
Romania. Herculane (, Gârnic (
Russia. Krasnodar Territory: Krasnodar(, Kuban distr (SB); Altai Krai: Zmeinogorsk (Obydov, 2010: 62); Zelenskaja Shehel riv. valley (SB); Daghestan: Magaramkentsky; Volgograd region: Olkhovskiy district (, Kamyshinsky district (; Kaliningrad region: Svetlogorsk, Sambian peninsula, Znamensk (Alekseev, 2008: 157); Saratov region: Saratov distr., Rtishchevskaya distr., Popovka distr., Voskresenskogo distr. (Sazhnev, 2007: 350); Moscow, Kazan, Tomsk (Breuning, 1927: 174); Tyumen Obl: Nizhnetavdinsky Distr, Sladkovsky Distr, Ishimsky Distr, Tyumen Distr, Ishim City, Omutinsky Distr, Armizonsky Distr, Kazansky Distr, Uporovsky Distr, Nizhnetavdinsky Distr, Vagaysky Distr; Omsk Obl: Bolsheukovsky Distr (, Gorkovsky Distr, Cherlaksky Distr Omsk City; Novosibirsk Obl: Karasuksky Distr; Kurgan Obl: Polovinsky Distr; Altai Krai: Zmeinogorsky Distr. (Stolbov & al, 2018: 733)
Serbia. Požarevac (Apfelbeck, 1904 : 16), Novi Sad (; Beli Potok (; Valjevo: Lelić (
Slovakia. Košice (Kassa) (Narozsny, 1938: 75)
Slovenia. Skrilje, Tolmin (Tolmino), Komen (Comeno), Mt. Slavnik (Taiano), Mt Snežnik (Nevoso) (Müller, 1926: 44); Prekmurje region (
Spain. Andalusia: Sevilla (Navarro & al., 2008); Avila: La Sefrada (Jeanne, 1969: 102); Balearic islands: Menorca, San Cristobal (Jeanne, 1969: 102); Barcelona: Barcelona, Castelldefels, Osona, Gavà, Begues, Vallgorguina, Garraf, Hostalets de Balenyà (, Tarrasa, Monteseny, Santa Fe (Jeanne, 1969: 102); Cádiz: San Roque, Algeciras (Durán, 1965: 66); Galicia: Pinol, Vilar Dastrés, Bordóns (Campos Gómez & al., 2006: 52); Girona: Romanyà, Gombrèn, Viladrau (, Nuria, Guillerias (Jeanne, 1969: 102), Huesca (SB), Sierra de Cazorla (; Granada: Jubiles (Jeanne, 1969: 102); Jaén: Santiago de la Espada (; Lérida: Coll d’Ager (Jeanne, 1969: 102); Leon: Quintanilla, La Laguna (Jeanne, 1969: 102); Lleida: Senterada, La Noguera (; Madrid: El Pardo (Jeanne, 1969: 102); Navarra: Villatuerta (, Ayegui (; Ourense: Vilar de Astrés, Carballeda de Valdeorras, Peña Trevinca (; Pontevedra: Sanxenxo, Tomiño, Sober (; Salamanca: Casillas de Flores, Guijo de Avila, Peña de Francia, Robleda (Zaballos, 1986: 75), Matilla, Buenamadre (Jeanne, 1969: 102); Tarragona: Fontscaldetes (
Syria. al Lādhiqīyah, Slinfah, 1130 m (SB), Lattakia ( 21033234)
Sweden. (Lindroth & Bangsholt, 1985: 47) Öland; Gotland (Breuning, 1927: 174); Dalarna Co.; Skåne: Falsterbo (
Switzerland. Zürich, Basel, Geneva, Thunersee, Canton of Valais, Aigle (Heer, 1838: 32), Chiasso (Magistretti, 1965: 58); Brugg distr: Bözen (
Tajikistan. Turkestan and partly Alai ranges, Zarafshan valley, Hissar-Darwaz (Mikhailov,; Verkhnie Khonaka valley (
Tunisia. région de Ouchteta (Bedel, 1895: 19), Jendouba, Ain Draham (SB)
Turkey. Anatolia (type of severum ; MNHN); Adalia: Thermessos (, Korkuteli (Tezcan & al., 2018: 549); Adana: Pozanti (Mandl 1963: 46 sub ab. purpureoaureum; SB); Amasia (type of rapax Motschoulsky, 1865: 309), Ankara (Mandl 1963: 46); Antalya: Yukarikaraman (, Güllük Dağı (Tezcan & al., 2018: 549); Ayaş (Kocatepe & Mergen, 2004: 297); Artvin: Hatila National Park, (Gokturk & Celik, 2017: 985); Balıkesir: Bigadiç, Doğanlar (Tezcan & al., 2018: 549); Bingol: Buglan Gecidi 1500 m (EM, SB); Bursa (Tezcan & al., 2018: 549); Çanakkale: Çanakkale (Kanat & Özbolat 2006), Lapseki (Tezcan & al., 2018: 549); Istanbul: Balökesir, Büyükada (Kanat & Özbolat 2006); Izmir: Izmir (Breuning, 1927: 173), Urla, Ödemiş, Bornova (Tezcan & al., 2018: 549); Karaman: Karadag, 1800m. ( ); Kahramanmaraş (Kanat & Özbolat 2006); Kocaeli: Gölcük (Tezcan & al., 2018: 549); Konya: Tuz Golu (EM, SB), Konya (; Manisa (Tezcan & al., 2018: 549); Mersin: Aydicink (SB), Erdemli (; Muğla: Mugla (SB), Fethiye Uzumlu (SB); Osmaniye prov : Hasanbeyli (Mandl 1963: 46); Samsun (Tezcan & al., 2018: 549); Şirnak: Andaç 1070m., (SB); Tunceli (Tezcan & al., 2018: 549); Uşak (Tezcan & al., 2018: 549); Van Prov.: Qatak Narli 1800m (Cesa News Nr 53)
Turkmenistan. Kopetdagh (Kryzhanovsky & Atamuradov, 1994: 416)
Ucraina. Mukacheve (Munkács), Berehovo (Beregszász) (Narozsny, 1938: 75)
United Kingdom. Devon: Darthmouth (Stephens, 1839: 16); East Sussex: Hastings (Stephens, 1839: 16), Eastbourne (; Leicestershire: Vale of Belvoir (; Norfolk: Yarmouth (Stephens, 1839: 16), Brighton (Courtis, 1848: 2113); Suffolk: Aldboro’ (Aldeburgh?) (Stephens, 1839: 16)
United States. Connecticut: Portland (, Cockaponset State Forest, Chester (UASM), New London co. (Ferreira, 2008: 307); Illinois: Chicago (; Delaware (Bousquet, 2012: 230); Maine: Lincoln Co. (; Maryland (Bousquet, 2012: 230), Chevy Chase (; Massachussets: Essex co. (UASM), Suffolk co., Boston (, Norfolk co. (; New Hampshire: Merrimack co., Strafford co. (; New Jersey: Morris co., Pine Brook (, West Milford, Manchester, Pine Brook (; Pennsylvania: Philadelphia co. (, Pike co. (; Rhode Island: Kent co. (; Virginia: Winchester co., Bull Run Mountains, Prince William co., Prince William Forest Park (Evans, 2009); Vermont; West Virginia (Bousquet, 2012: 230).
Uzbekistan. Margilan (sub var. purpuripennis; Reitter, 1891: 257); Khorezm reg., Karakapakstan rep. (Khamraev, 2003: 47)

Notes: Winged diurnal but in some case can be attracted to light at night. It is inhabitant of both coniferous and decidous forests. Adults and larvae are excellent climbers and feed on Lymantridae and Thaumatopoeidae (Thaumatopoea pityocampa Denis & Schiffermuller, Lophirus pini Linné, Thaumetopoea processionea Linné, Lymantria dispar L., Euproctis chrysorrhoea Linné) and other caterpillars infesting trees of genera Pinus, Quercus and Fagus. Adults hunt trunks and treetops during the day and go down to the ground at around sunset and then hide in leaf litter at the foot of the trees.
Captures of active individuals have been noted from April to August and they are quite long lived as they seem able to survive up to 4 years (Dowden, 1962: 51). It is not rare to find specimens overwintering in small cavities in the ground.
Calosoma sycophanta is an handsome beetle that has attracted, since the first steps of entomology, the interest of entomologists. It is one of the few Carabidae represented in the volume of Aldovrandi (De Animalibus Insectis Libri septem cum singulorum Iconibus ad vivum expressis 1638: 450, fig. 6) under the name of Scarabaeus viridis. Later Reamour (Memoires pour servir a l'histoire des Insectes, vol. II, 1736: 455 and plate 37 fig. 18) has given an account of its way of life. Subsequently there have been numerous illustrations in books of this beautiful species. It can still be interesting to recall that, because of its rarity in the British Isles, Donovan (The Natural History Of British Insects, vol. XIV, 1810: pl.477) has drawn in its place the Calosoma (Calodrepa) scrutator. This error has been then corrected by Curtis (British Entomology, 1823-1840: pl.330).
The larva of C. sycophanta has been repeatedly described and drawn, starting with the first news given by Reamour (1736, vol. II: plate 37 fig. 14, 17).

Reamour Memoires pour servir a l'histoire des Insectes, vol. II, 1736: 455 and plate 37 fig. 18
Aldovrandi (De Animalibus Insectis Libri septem cum singulorum Iconibus ad vivum expressis 1638: 450, fig. 6) under the name of Scarabaeus viridis
Calosoma (Calosoma) sycophanta
(Linné, 1758)
Italia, dint. Roma, Manziana, I.72,, Bruschi lgt.
Calosoma (Calosoma) sycophanta
(Linné, 1758)
Turkey, Pozanti, 30.4.09, Boilly
Calosoma (Calosoma) sycophanta
(Linné, 1758)
Italia, Lazio, Bagnoregio, VII.1974, Bruschi lgt.
Calosoma (Calosoma) sycophanta
(Linné, 1758)
Moravia (Calosoma sycophanta azureum det. Mandl)
(coll. Naturhistorischen Museum Basel)
Calosoma (Calosoma) sycophanta
(Linné, 1758)
Elbourz, Talyche, de 0 à 2500m, J. de Morgan
(type of Calosoma sycophanta ssp. prasinum Lapouge, 1907)
(coll. Muséum National d'Histoire Naturelle, Paris)
Calosoma (Calosoma) sycophanta
(Linné, 1758)
Iran, Gilan prov, Rasht distr., near Dizkohv vill.
5-6.VI.2016, leg. Kasatkin D. lgt
Calosoma (Calosoma) sycophanta
(Linné, 1758)
Sardegna, Casa Cantoniera di Bidicolai, SS 125 Orientale Sarda, VIII.2018, remains found by M. Zilli
Calosoma (Calosoma) sycophanta
(Linné, 1758)
(type of Calosoma sycophanta var. severum Chaudoir, 1850)
(coll. Muséum National d'Histoire Naturelle, Paris)
Calosoma (Calosoma) sycophanta
(Linné, 1758)
Turkey, Pozanti, 30.4.09, Boilly
Calosoma (Calosoma) sycophanta
(Linné, 1758)
Turkey, Pozanti, 30.4.09, Boilly
updated April 3 2020