Calosoma (Camedula) angulatum Chevrolat, 1834

Length 25-33 mm. C. angulatum is characterized by very angular pronotum and deep striation of elytra, on which base was created Carabosoma subgenus, that later Jeannel included inside of Camedula.
The shape of pronotum and the sculpture of elytra vary quite regularly from the northern to the most southerly populations: the populations of Colombia and Venezuela, in particular, have relatively less angular pronotum, that is also more densely dotted, and more scaly intervals.
Based on these characteristics, Jeannel has dealt with these populations as one separate species (angulicolle Chaud. 1869). Vice versa, Gidaspow (1959: 252, 1963: 280) has considered conspecific, angulatum and angulicolle, as later confirmed by Erwin (1991: 26).
C. angulatum, thus interpreted, is present in Arizona and occasionally in California, Texas and New Mexico. It is widespread in Mexico, and reappears in Central America. It has been found in Costa Rica, Nicaragua, Guatemala, El Salvador and Honduras. It is present in Colombia and, according to old citations, also in Venezuela (Breuning, 1928a: 103)

Examined specimens and literature’s data
Colombia. Santa Marta: Santa Marta (MNHN), Bonda (AMNH); La Guajira: Uribia, Macuira PNN (N. Forero i.l.)
Costa Rica. La Caja, San José (Erwin: 1991: 28); Puntarenas: Lepanto, Monteverde (www.gbif.org/); Guanacaste: Santa Elena (www.gbif.org/), Santa Rosa (Erwin: 1991: 28), Tronadora, Liberia (www.inaturalist.org)
El Salvador. Santa Ana: Metapán (www.inaturalist.org/obs/56087932)
Guatemala. Zacapa: San Jorge (www.inaturalist.org/obs/14732178)
Honduras. El Zamorano (www.inaturalist.org/obs/50939261)
Mexico. Aguascalientes: Aguascalientes (www.inaturalist.org); Baja California: Guadalupe Island (Erwin, 2007: 86); Chiapas: Villa Morelos, Comitan, Junction Highways 190-195 (UASM); Chihuahua (Bates, 1884: 262); Colima: Tecoman, Colima (UASM), Manzanillo (www.inaturalist.org); Guanajuato: Palo Verde (Bates, 1884: 20), Irapuato (www.inaturalist.org); Guerrero: Ciudad Altamirano (SB), Zihuatanejo de Azueta, Tixtla de Guerrero (www.inaturalist.org); Jalisco: Bahia de Cuastecomate (UASM), Guadalajara, Melaque (SB), Zapopan, Ahualulco de Mercado, Autlan de Navarro, La Huerta, Rancho Vazquez, El Grullo, Chapala (www.inaturalist.org/); Michaocan: La Placita (AVT), Tiquicheo (SB), Huetamo (www.entomologiitaliani.net), Nocupétaro (www.inaturalist.org/); Morelos: Tejalpa (UASM), Cuernavaca (JM), Cuautla (www.inaturalist.org/); Nayarit: Ahuacatlan (UASM), La Yesca, Tepic, Puerto Balleto (Isla María Madre) (www.inaturalist.org/); Nuevo Leon: Lampazos de Naranjo, General Escobedo, San Pedro Garza García, Monterrey (www.inaturalist.org/); Oaxaca: Tehuantepec (Bates, 1884: 21), Juchatengo (UASM), Santiago Astata (http://www.ebay /), P.N. Huatulco (www.inaturalist.org/); Puebla: Esperanza (Bates, 1884: 21), Zacapala, Cholula de Rivadabia, Izúcar de Matamoros (www.inaturalist.org/); San Luis Potosi: Tamuin (SB), Ciudad Valles, El Naranjo (UASM); Sinaloa: La Noria, Guasave, Culiacan, Mazatlan (UASM); Sonora: Aconchi, Moctezuma, Yecora, Alamos, Sierra de Alamos, Mazatan, Bacanora Nacozari, Tecoripa, (UASM); Tamaulipas: La Mesa, Rio Sabinas, Magiscatzin, Ciudad Mante, Villa Manuel, Villa de Casas (UASM), Ciudad Victoria, Llera (www.inaturalist.org); Veracruz: Plan del Rio (Bates, 1884: 21), Boca del Monte (type C. angulatum MNHN), Palma Sola (UASM)
Nicaragua. Granada: Reserva Natural Vocan Mombacho (www.gbif.org/); León (www.gbif.org/); Chinandega (www.bio-nica.info/); Reserva Ecologica El Bajo (www.inaturalist.org/); Carazo (Su et al., 2005: 145)
United States. Arizona: S. Cruz County (EM, SB), Santa Rita Experimental Station, Santa Rita Mountains (www.gbif.org/), Pima County (http://bugguide.net/), Tucson, Cochise county (www.inaturalist.org); California (Burgess & Collins, 1917: 75); New Mexico (Gidaspow, 1959: 253); Texas: Cameron County (http://bugguide.net/)

Notes: Winged, often attracted to light at night. It lives from lowlands up to an altitude of 1800m. In Nicaragua it has been reported feeding on the noctuid caterpillar Spatodoptera frugiperda. Active adults have been noted from May to November (Erwin, 2007: 86).

Calosoma (Camedula) angulatum ♂ Chevrolat, 1834 United States, Arizona: S. Cruz County, Rio Rico, VII.75 Lenczy leg.

Calosoma (Camedula) angulatum ♀ Chevrolat, 1834 United States, Arizona: S. Cruz County, Rio Rico, VII.75 Lenczy leg.

Calosoma (Camedula) angulatum ♂ Chevrolat, 1834 Mexico,Morelos, Cuernavaca 23.06.2011 (photo and coll. J. Matějíček)

Calosoma (Camedula) angulatum ♀ Chevrolat, 1834 Nouvelle Grenade ou Venezuela (Chili?) (type of Calosoma angulicolle Chaudoir, 1869) (coll. Muséum National d'Histoire Naturelle, Paris)