Calosoma (Caminara) davidis Géhin, 1885

Calosoma lugens var. Davidis Géhin, 1885: 61, note 61 (described from: Mou-Pin); original material unspecified; type in former coll. Oberthur (Jeannel, 1940: 119)
Calosoma thibetanum Farmaire, 1887: 92 (type locality: Mou Pin); holotype ♀ by monotypy designated by Deuve (1978: 248) in Muséum National d'Histoire Naturelle, Paris [examined]
Calosoma (Caminara) davidis, Breuning, 1927: 226
Calosoma (Caminara) arrowianum Breuning, 1934b: 48 (type locality: Distrikt Zayul, Atakang 2700m.); holotype ♂ in British Museum of Natural History, London (Jeannel, 1940: 119)
Catasoma thibetanum, Lapouge, 1930: 98
Campalita davidi, Jeannel, 1940: 119
Campalita arrowianum, Jeannel, 1940: 119
Calosoma (Caminaria) davidis martensi Paulus, 1971: 16 (type locality: W. Nepal, Jomosom 2750m.) type material: holotype ♂ and 10 paratypes, depository not stated
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Length 22-28 mm., C. davidis has a large, transverse, pronotum with sligthly angular sides. Its elytral sculpture is of "pentaploïde" type as it has two tertiary intervals on each sides of a secondary one and it is of "hétérodyname" type, as the intervals are not equal in width and height. In fact the primary intervals of C. davidis consist in series of chain-like elevations with opaque foveae of the same dark color as the rest of elytra, the secondary intervals, just raised but still visible, are represented by series of aligned grains, while the tertiary ones are transformed into disorderly graininess.
From Nepal has been described "Calosoma (Caminaria) davidis martensi" (Paulus, 1971:16), with elithra slightly more elongated and parallel, and with slightly more transverse pronotum. However these characters dont appear to justify a taxonomic separation between the populations of Nepal and China.
Also Calosoma (Caminara) arrowianum (Breuning, 1934), which was described on a single specimen from southern Thibet "Zayul district, Atakang" (= Zayü county), near the Indian border, has to be referred to C. davidis. In fact, considering the scarce significance of the characters reported in the original description and comparing the numerous recent specimens coming from the same area, the specific distinction proposed between davidis and arrowianum appears unnecessary. Jeannel too (1940: 119), reporting the original description, without having seen the type, considered morphologically very close the two supposed species.
C. davidis is found in western China, including southern Thibet, in the neighboring Nepal and northern Myanmar.

Examined specimens and literature’s data
China. Hubei: Yichang (Deve, 1997: 55), Jinjenshan, 1500 m., VI.93 1 ♂ (AVT); Shaanxi: Tai Bai Shan (AVT, SB); Sichuan: Jin-fu Shan (EM, AVT), Xichang (SB), Shimian (SB, EM); Xizang (= Tibet): Mou Pin (type thibetanum MNHN); Linzhi city (Nyingchi Pref.) (SB), Chayu (= Zayü county) (SB), Zayü county, Ata Kang La, (type arrowianum, Jeannel, 1940: 119), Yigongxiang (Bomê co.) (Deuve, 1997: 55); Yunnan: Pe Ye Tsing (= Yanfeng) (SB), Zhaotong (SB), Dali (Tali) (Jeannel, 1940: 119)
Myanmar. Kachin, Chudo Razi 2800m (SB)
Nepal. Thakkhola (= Thak Khola, upper part of the Kali Gandaki valley) Jomsom (= Jomsom) 2750m. (type of martensi, Paulus, 1971: 20); Kali Gandaki valley: Kagbeni 2800m (JS), Marpha 2650m (JS); C. Nepal, Dzhonston (?) (SB).

Notes: Winged. Active individuals were captured from May to July, during the raining season. The Nepalese specimens were captured in cultivated fields using fall traps (Paulus, 1971: 20)
We must note that the species has been described as C. Davidis without any explicit dedication. Starting from Jeannel (1940) it has been sometimes referred to as C. Davidi. The name given by the author is properly formed in accordance with the rules of the International Code of Zoological Nomenclature (art 31.1.1.), corresponding apparently to the genitive case of the Latin name of the biblical hero David, and the prevailing usage rule (art. 23.9.1) does not seem applicable in this case. Therefore C. davidis is the only name that must be used.

Calosoma (Caminara) davidis
Géhin, 1885
Central Nepal, Dzhonston, 9.5.96, Cherniakovsky lgt.
Calosoma (Caminara) davidis
Géhin, 1885
Central Nepal, Dzhonston, 9.5.96, Cherniakovsky lgt.
Calosoma (Caminara) davidis
Géhin, 1885
China: Yunnan, Pe Ye Tsing
Calosoma (Caminara) davidis
Géhin, 1885
China: Yunnan, Pe Ye Tsing
Calosoma (Caminara) davidis
Géhin, 1885
Mou Pin, A David
(holotype of Calosoma thibetanum Farmaire, 1887)
(coll. Muséum National d'Histoire Naturelle, Paris)
last review April 23 2019

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