Calosoma (Caminara) denticolle Gebler, 1833

Calosoma denticolle Gebler, 1833: 274 (Typus: Loktevsk; Zoological Institute, Russian Academy of Sciences, St. Petersburg)
Calosoma granulosum Motschulsky, 1844: 121
Calosoma lugubre Motschulsky, 1844: 122
Calosoma rugulosum Motschulsky, 1850: 89
Callistrata granulosum Motschulsky, 1865: 306
Callistrata denticollis androgyna Lapouge, 1930: 90
Callistrata denticollis mongolica Lapouge, 1930: 91 (described from: Mongolia)
Charmosta (Callistrata) denticolle granulosa Lapouge, 1932: 424
Calosoma (Charmosta) denticolle Breuning, 1927: 229
Campalita denticolle Jeannel, 1940: 120
Caminara denticolle Basilewski, 1972: 37
Calosoma (Charmosta) denticolle suensoni Mandl, 1981: 21 (Typus: Yüncheng; Zoologischen Museums der Universität, Kopenhagen)

C. denticolle has two tertiary intervals on each sides of a secondary one ("pentaploïde" type). The intervals, that are deeply incised, are all of the same width and height ("homodyname" type), and all of them are transformed into series of rough granules, more or less aligned. C. denticolle can be easily distinguished from the other species collected in Caminara by the acute basal angles of the pronotum, by the body shape stout and convex and by the upper body with noticeable bronzed reflexes with large coppery metallic foveae on the primary intervals, and finally by the male anterior tarsi not dilated.
It occupies a large area, from Black sea western coast to almost the Baltic sea and trought southern Russia, Crimea, Caucasus, Iran, Central Asia, southern Siberia, Mongolia, North China. The northernmost records consist in some few specimens found along the southern sea coast of Finland, possibly dragged by strong gales in such an unsuitable environment.

Examined specimens and literature’s data
Armenia: Yerevan, (Breuning, 1927: 229)
Belarus: Turov env., Arevithcy vill., Polesie State Radioecological Reserve (Aleksandrowicz, 2011: 478)
Bulgaria: Southern Black Sea coast: Arkutino, Sozopol (Guéorguiev & Guéorguiev, 1995: 41); Nesebar (http://www.ebay.it)
China. Liaoning: Xiuyan (SB); Shaanxi: Yüncheng (paratype of suensoni) (NMB); Heilongjiang; Ningxia Dawukou (Deuve, 1997: 56), Mt.Old father, Mudanjiang (SB)
Finland: Tvärminne, Brännskä (Lindroth & Bangsholt, 1985: 49)
Georgia: Abkhazia, (Breuning, 1927: 229)
Iran: Golestan Gonbad (Ghahari et Al 2009: 439)
Kazakhstan: Aksaj Karačaganak (SB); Dzanybek (SB); Semipalatinsk (=Semej) (SB); Borochoro chain (Breuning, 1927: 229)
Moldova: Southern Moldova, right bank of Dnestr river (Neculiseanu et Al, 2000: 40)
Mongolia: 50 km N Ulaanbaatar (= Ulan Bator) (SB); 25 km. NE Tsejserteg, Arkhangay (SB); 100 km. SSW Baruun-Urt (SB)
Romania: Mangalia (SB)
Russia: Volgograd distr (SB); southern Russia (EM); Kalmykia reg. (SB); Sazara prov. (SB); Buren (near the Mongolian border) (SB); Barnaul (SB); Sotschi (VV); Rjasan; Viatka (Breuning, 1927: 229); Daghestan (Breuning, 1927: 229); Bouriazia, Kiakhta (Jeannel, 1940: 120); Tomsk (Breuning, 1927: 229)
Turkey: near Istanbul (Jeannel, 1940: 120)
Turkmenistan (Kryzhanovsky & Atamuradov, 1994: 416)
Ucraine: Sevastopol (Jeannel, 1940: 120); Poltawa (Breuning, 1927: 229); Kherson (SB)
Uzbekistan: Khorezm reg., Karakapakstan rep. (Khamraev, 2003: 47)

Notes: Winged. Adults are active from June to August (according to the material examined) and in September (Mandl, 1981: 22) and they can be captured in pit traps.

Calosoma (Caminara) denticolle
Gebler, 1833
Romania: Mangalia, 14-16.8.90, T. Tračka lgt.
Calosoma (Caminara) denticolle
Gebler, 1833
Kazakhstan: Aksaj Karačaganak, VI.88, Čermàk lgt.
Calosoma (Caminara) denticolle
Gebler, 1833
China: Yüncheng, 12.IX.1929 Suenson lgt. (paratype of
Calosoma (Charmosta) denticolle suensoni Mandl, 1981)
(coll. Naturhistorischen Museum Basel)

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