Calosoma (Caminara) denticolle Gebler, 1833

Calosoma denticolle Gebler, 1833: 274 (type locality: near Loktevsk); original material: 2 ♀♀; type in Zoological Institute, Russian Academy of Sciences, St. Petersburg (Jeannel, 1940: 120)
Calosoma granulosum Motschulsky, 1844: 121 (described from: Lac Inderskoie) type material not stated and no repository given
Calosoma lugubre Motschulsky, 1844: 122 (described from: Turkménie) type material not stated and no repository given
Calosoma rugulosum Motschulsky, 1850: 89 (described from: Kharkov)
Callistrata granulosum Motschulsky, 1865: 306
Calosoma (Charmosta) denticolle Breuning, 1927: 229
Callistrata denticollis androgyna Lapouge, 1930: 90 (distribution: de la Prusse à la Mandchourie) type material not stated and no repository given
Callistrata denticollis subsp mongolica Lapouge, 1930: 91 (distribution: depuis l'Altai jusq'en Mongolie) type material not stated and no repository given
Charmosta (Callistrata) denticolle granulosa Lapouge, 1931: 424
Campalita denticolle Jeannel, 1940: 120
Caminara denticolle Basilewski, 1972: 37
Calosoma (Charmosta) denticolle ssp. suensoni Mandl, 1981: 21 (type locality: Yüncheng); holotype ♂ and paratypes in Zoologischen Museums der Universität, Kopenhagen and in Naturhistorischen Museum Basel [examined]

C. denticolle has two tertiary intervals on each sides of a secondary one ("pentaploïde" type). The intervals, that are deeply incised, are all of the same width and height ("homodyname" type), and all of them are transformed into series of rough granules, more or less aligned. C. denticolle can be easily distinguished from the other species collected in Caminara by the acute basal angles of the pronotum, by the body shape stout and convex and by the upper body with noticeable bronzed reflexes with large coppery metallic foveae on the primary intervals, and finally by the male anterior tarsi not dilated.
C. denticolle suensoni, from the Chinese Southern region of Guangdong, should be characterized by the roughness of head and pronotum and by the shape of the pronotum that should be more shorter and wider, narrowed to the base with more pointed posterior angles. These characters are very variable in the specimens of the entire area of diffusion and the elevation of the population of Yuncheng to the rank of subspecies appears superfluous.
C. denticolle occupies a large area, from the Baltic sea to Black sea, and from the western coast of the Black sea to Far East trought southern Russia, Caucasus, northern Iran, Central Asia, southern Siberia, Mongolia, North China. The northernmost records consist in some few specimens found along the southern sea coast of Finland, possibly dragged by strong gales in such an unsuitable environment.

Examined specimens and literature’s data
Armenia. Yerevan, (Breuning, 1927: 229)
Belarus. Turov env., Arevithcy vill., Polesie State Radioecological Reserve (Aleksandrowicz, 2011: 478)
Bulgaria. Southern Black Sea coast: Arkutino, Sozopol (Guéorguiev & Guéorguiev, 1995: 41); Nesebar (http://www.ebay.it)
China. Guangdong: Yüncheng (paratype of denticolle suensoni) (NMB); Heilongjiang: Mt.Old father, Mudanjiang (SB); Liaoning: Xiuyan (SB); Ningxia: Dawukou (Deuve, 1997: 56); Sichuan: Hanyuan (Ebay, 2014)
Finland. Tvärminne, Brännskär (Lindroth & Bangsholt, 1985: 49)
Georgia. Abkhazia, (Breuning, 1927: 229)
Iran. Golestan: Gonbad (Ghahari et Al 2009: 439)
Kazakhstan. Aksaj Karačaganak (SB); Dzanybek (SB); Semipalatinsk (=Semej) (SB); Almaty: Kashengel (SB); Borochoro chain (Breuning, 1927: 229)
Moldova. Southern Moldova, right bank of Dnestr river (Neculiseanu et Al, 2000: 40)
Mongolia. 50 km N Ulaanbaatar (= Ulan Bator) (SB); 25 km. NE Tsejserteg, Arkhangay (SB); 100 km. SSW Baruun-Urt (SB), Domogov reg., 2km SE Khuvsgol (SB)
Romania. Mangalia (SB)
Russia. Altai Krai: Loktevsky District (Gebler, 1833: 274); Volgograd distr (SB); Olkhovskiy distr (http://molbiol.ru/); southern Russia (EM); Kalmykia reg. (SB); Sazara prov. (SB); Buren (near the Mongolian border) (SB); Barnaul (SB); Sotschi (VV); Saratov reg: Saratov distr., Balashov distr., Krasnoarmeyskiy distr., Rtischevsky distr. (Sazhnev, 2007: 351); Orenburg reg: Akbulak distr (SB); Krasnodar reg.; St. Petersburg env. (SB); Rjasan; Viatka (Breuning, 1927: 229); Daghestan (Breuning, 1927: 229), Gergebil'skij distr. (www.gbif.org); Bouriazia: Kiakhta (Jeannel, 1940: 120); Tomsk (Breuning, 1927: 229), Astrakhan reg. (SB); Čeljabinsk reg: Kizel'skij distr. (www.gbif.org)
Turkey. near Istanbul (Jeannel, 1940: 120)
Turkmenistan. (Kryzhanovsky & Atamuradov, 1994: 416); Ahal prov. (www.gbif.org)
Ucraine. Sevastopol (Jeannel, 1940: 120); Poltawa (Breuning, 1927: 229), Kherson (SB)
Uzbekistan. Khorezm reg., Karakapakstan rep. (Khamraev, 2003: 47)

Notes: Winged. Adults are mainly active from May to September (according to the material examined) and they can be captured in pit traps.

Calosoma (Caminara) denticolle
Gebler, 1833
Romania: Mangalia, 14-16.8.90, T. Tračka lgt.
Calosoma (Caminara) denticolle
Gebler, 1833
Kazakhstan: Aksaj Karačaganak, VI.88, Čermàk lgt.
Calosoma (Caminara) denticolle
Gebler, 1833
China: Yüncheng, 12.IX.1929 Suenson lgt. (paratype of
Calosoma (Charmosta) denticolle suensoni Mandl, 1981)
(coll. Naturhistorischen Museum Basel)
last review April 23 2019

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