Calosoma (Carabomimus) depressicolle Chaudoir, 1869

Calosoma depressicolle Chaudoir, 1869: 375 (type: Toluca; coll. Oberthur, Muséum National d'Histoire Naturelle, Paris)
Calosoma (Blaptosoma) depressicolle Breuning, 1928: 54
Blaptosoma (Eutelodontum) depressicolle Lapouge, 1932: 392
Carabomimus depressicollis Jeannel, 1940: 229
Carabomimus depressicollis asper Jeannel, 1940: 229 (type: Mexico; presently not to be found)
Carabomimus depressicollis altipeta Jeannel, 1940: 229 (lectotype: volcan du Nevada; Muséum National d'Histoire Naturelle, Paris)
Calosoma (Carabomimus) depressicolle Gidaspow, 1959: 296
Calosoma (Carabomimus) asper Gidaspow, 1959: 297
Calosoma (Carabomimus) altipeta Gidaspow, 1959: 297
Carabomimus depressicollis silvestris Lassalle, 2009: 3 (type: Rio Frio; coll. Lassalle)
Calosoma (Carabomimus) depressicolle asper Häckel et Al., 2011: 48.
Calosoma (Carabomimus) altipeta Häckel et Al., 2011: 51.
Calosoma (Carabomimus) silvestre Häckel et Al., 2011: 51.


Length 18-22 mm. C. depressicolle, C. flohri and C. digueti, are characterized by the slightly heart shaped pronotum with evident hind angles.
C. depressicolle and C. digueti both have a smooth pronotum without punctures, but C. depressicolle is further distinguished because of the apex of its penis simple, without any apophysis.
All the populations of C. depressicolle have the same shape of penis and similar characteristics of the sculpture of elytra, with evident and confused intervals. The dimensions are somewhat greater in eastern populations.
Jeannel (1940) on the basis of a comparison with the type from Toluca, that is small, with narrower pronotum and less elevated sculpture of elytra, described, from the surroundings of Ciudad de Mexico, a population (C. depressicollis asper) distinguished by the larger size, slightly wider pronotum and less regular sculpture of elytra, with raised intervals.
Another entity that Jeannel considered as subspecies of C. depressicollis was Calosoma (Blaptosoma) lesnei (Breuning, 1931: 54) that we consider to be a synonymous with C. flohri, and which we will deal with treating of this last species.
From the mountain near Toluca but on higher altitude, Jeannel (1940) described another population (C. depressicollis altipeta) of smaller dimension and more stout body shape, with sculpture of elytra always strong enough, but with sharper apex of the penis.
Gidaspow (1959), based on the descriptions of Jeannel, but according to the criterion of sympatry, or contiguity of areal, has considered these populations as distinct species.
Lassalle & van den Berghe (2011), in a general revision of the group, came back to the interpretation of Jeannel, dealing with all these populations as subspecies, including the population of Rio Frio (Mexico) that Lassalle (2009) had previously described as C. depressicollis silvestris, taking into account an average larger size and more confused sculpture of elytra.
On the other hand, Häckel & al. (2011), accepted C. depressicollis asper at subspecies level, but have considered altipeta and silvestre, as distinct species, even if collected together in the same group of species.
In any way, we think that geographical considerations, together with the inconsistency of morphological differences, should lead to collect all these populations within C. depressicolle, remaining separated from C. flohri, by the differences of the aedeagus. The various populations of C. depressicolle, are found in the state of Mexico, and penetrate, just beyond the border states of Michoacán (Zitacuaro, Häckel & al., 2011) and Morelos (Huitzilac, Breuning, 1928b: 55). There are also quotes for the south of the state of Jalisco but these, after the most recent reviews of the group (Lassalle, 2009; Lassalle & van den Berghe: 2011), should be verified because most likely they should be referred to C. digueti colimaensis

Examined specimens and literature’s data
Mexico. Distrito Federal: El Guardia, 3000 m (EM, SB), Salazar 3050 m (GP), Volcán Ajusco (Häckel et Al., 2011: 51); Mexico: Rio Frio, 3200 m (AVT, EM, SB), Santa Marta, Ocuilan 2700 m (EM, SB), Cañada de Alferes 2500 m (GP), Volcan du Nevada (lectotype of altipeta, MNHN), Nevado de Toluca (GP), Toluca (holotype of depressicolle MNHN), between Toluca and Nevado de Toluca 3200 m (EM, SB), Vale de Bravo, Villa de Allende, Coatepec (Häckel et Al., 2011: 51), Tenancingo, Amanalco, Bosencheve National Park, San Cayetano, W of Toluca N of Agua Bendita, Amecameca, La Hortaliza 43 km W of Toluca, 6.6 km E of Rio Frio (UASM); Michoacan: Zitacuaro (Häckel et Al., 2011: 51); Morelos: Huitzilac (Breuning, 1928b: 55)

Notes: Brachypterous. It is nocturnal, and lives in woodlands, or in their fringe, where it is active in April-May and July-September, or, according to Erwin (2007: 88), up to November.

Calosoma (Carabomimus) depressicolle
Chaudoir, 1869
Mexico: between Ciudad de Mexico and Tuxmelucan,
Rio Frio, m 3200, 16.V.1980, S. Bruschi leg.
Calosoma (Carabomimus) depressicolle
Chaudoir, 1869
Mexico: between Ciudad de Mexico and Tuxmelucan,
Rio Frio, m 3200, 16.V.1980, S. Bruschi leg.
Calosoma (Carabomimus) depressicolle
Chaudoir, 1869
Mexico: Toluca, Volcan de Nevado, 3200m, L Diguet.
(lectotype de Carabomimus depressicollis altipeta Jeannel)
(coll. Muséum National d'Histoire Naturelle, Paris)
Calosoma (Carabomimus) depressicolle
Chaudoir, 1869
Mexico: between Huitzilac and Coatepec,
S. Marta, m 2700, 18.VII.1987, S. Bruschi leg.

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