Calosoma (Carabomimus) flohri Bates, 1884

Calosoma flohri Bates, 1884: 262 (lectotype: Uruapan; Muséum National d'Histoire Naturelle, Paris)
Calosoma (Blaptosoma) laeve var. gebieni Breuning, 1928: 46 (type: Pazcuaro; Biozentrum Grindel und Zoologisches Museum, Hamburg, presently not to be found)
Calosoma (Blaptosoma) cicatricosum ssp. flohri Breuning, 1928: 53
Calosoma (Blaptosoma) lesnei Breuning, 1931: 54 (holotype: Tlapujahua, Muséum national d’Histoire naturelle, Paris)
Blaptosoma (Eutelodontum) cicatricosum flohri Lapouge, 1932: 392
Carabomimus depressicollis flohri Jeannel, 1940: 229
Carabomimus gebieni Jeannel, 1940: 229
Carabomimus depressicollis lesnei Jeannel, 1940: 229
Calosoma (Carabomimus) gebieni Gidaspow, 1959: 289
Calosoma (Carabomimus) flohri Gidaspow, 1959: 295
Carabomimus flohri Lassalle, 2009: 8
Calosoma (Carabomimus) ortrudae Häckel & Heinz, 2011: 2 (type: 48km E Morelia 2900m.; coll. W Heinz)
Calosoma (Carabomimus) flohri gebieni Häckel et Al., 2011: 52.
Calosoma (Carabomimus) lesnei Häckel & al., 2011: 54.
Calosoma (Carabomimus) flohri zirostense Häckel et Al., 2011: 54.(type: Michoacan, env. Nuevo Zirosto 800m., Staatlichen Museum für Naturkunde Stuttgart, presently coll. W Heinz).
Carabomimus lesnei Lassalle & van den Berghe, 2011a: 50


Length 18-22 mm. C. flohri was regarded by Jeannel as a subspecies of C. depressicolle.
As C. depressicolle, C. flohri is characterized by heart-shaped pronotum but it has a less raised and more regular sculpture of elytra. Moreover it is distinct from C. depressicolle, which is found more south, because of the strong punctures on the head and pronotum and because of the apex of penis which has a characteristic angular apophysis projecting on the left edge.
C. flohri was described from Uruapan in the state of Michoacán. and it is widespread in the mountain ranges of this state. Gidaspow (1959: 296) also indicates Colima and near Toluca in Mexico state, and Erwin (2007: 92) adds Jalisco. The presence of C. flohri near Toluca is very unlikely and possibly comes from a misinterpretation of a quote by Bruning (see below), but but it cannot be excluded that C. flohri can also be found somewhere beyond the border in neighboring states. of Colima, Mexico, and Jalisco.
From Patzcuaro , a few kilometers east of the type locality, Breuning (1928b: 46) described Calosoma (Blaptosoma) laeve var. gebieni. It was revised and illustrated by Jeannel (1940) that stated it should belong to a distinct species. According to the diagnosis of Jeannel, C. gebieni has the same type of genital of C. flohri, its base of pronotum and head are dotted on the margins only, the and elytra are oblong, smooth and shiny. According to Lassalle (2009: 8), C. gebieni is a synonym of C. flohri, because the features, on which the description is based, are within the variability of this last species.
Later, Breuning (1931) described another population (C. depressicollis lesnei) from the Sierra of Tlapujahua on the border between Michoacán and Mexico and not near Toluca as erroneously stated by Jeannel (1940: 231) and Gidaspow (1959: 295). C. depressicollis lesnei is characterized by less raised sculpture of the elytra and by a less pronounced apophysis projecting on the apex of penis. Considering that the characters on which the description was based, including the shape of the apex of penis, appear to be identical or may fall within the variability of C. flohri Gidaspow (1959: 296) expressed the opinion that it should be a sinonimous to this last species, and, after having examined the type and the specimens above referred to, we share this interpretation. However, we have to add that Lassalle & van den Berghe (2011), in a general revision of the group, considered C. lesnei a separate species.
Also Calosoma (Carabomimus) ortrudae (Häckel & Heinz 2011: 2), described as distinct species and found between Morelia and Ciudad Hidalgo, considering the exterior morphological appearance and its localisation, is probably a population of C. flohri even if the descriptors have seen it as close to C. digueti.
As we prefer to avoid to use the concept of complex of species or to limit the recourse to infraspecific divisions, we collect under C. flohri all the populations that in some way can be referred to this species, whatever the level of the hierarchy proposed by the authors. Who will want, will decide freely on the systematic position of the various entities through direct consultation of the literature cited.

Examined specimens and literature’s data
Michoacán: Uruapan (lectotype of C. flohri, MNHN); Angahuán, S of Carapán (Häckel et Al., 2011: 52); Corupo, Tancitaro, cerro Taretzuruan (Gidaspow, 1959: 296); 10 km. S. Paracho 2340 m (EM); between Uruapan and Paracho 2200 m (EM, SB); San Juan Tumbio (SB); between Uruapán and Pátzcuaro (Häckel et Al., 2011: 52); Patzcuaro (holotype of gebieni, MNHN); between Cheran and Carapán, 6,2 miles W of Quiroga, Ojo de Agua Grande (sub flohri gebieni Häckel et Al., 2011: 52); Sierra of Tlapujahua (holotype of lesnei, MNHN), 15 km W Ciudad Hidalgo 2200m (SB); Laguna Verde (sub lesnei Häckel et Al., 2011: 55); Nuevo Zirosto (type of zirostense, Häckel et Al., 2011: 54); José de la Cumbre (type of ortrudae, Häckel & Heinz, 2011: 2)

Notes: Brachypterous. It is nocturnal, in daytime takes frequently refuge under stones or pieces of wood. Lives in woodlands, their fringe, radures, pastures and grasslands where it is active from May to August.
The name of the species should remember the one of Julius Flohr (1837–1896) a physician born in Germany, that, in 1859, moved to Mexico where he made extensive entomological collections.

Calosoma (Carabomimus) flohri
Bates, 1884
Michoacan: Uruapan (lectotype) (coll. Muséum National d'Histoire Naturelle, Paris)
Calosoma (Carabomimus) flohri
Bates, 1884
Michoacan:15 km west Ciudad Hidalgo,
m 2200, 10.VII.1987, S. Bruschi leg.
Calosoma (Carabomimus) flohri
Bates, 1884
Michoacan: between Uruapan and Paracho,
m 2200, 11.VII.1987, S. Bruschi leg.
Calosoma (Carabomimus) flohri
Bates, 1884
Michoacan: between Uruapan and Paracho,
m 2200, 11.VII.1987, S. Bruschi leg.
Calosoma (Carabomimus) flohri
Bates, 1884
Mexico: Sierra de Tlapujahua,
env. de Toluca 2400 2700m, L Diguet.
(type de Calosoma (Blaptosoma) lesnei Breuning)
(coll. Muséum National d'Histoire Naturelle, Paris)
Calosoma (Carabomimus) flohri
Bates, 1884
Mexico: Sierra de Tlapujahua,
env. de Toluca 2400 2700m, L Diguet.
(coll. Muséum National d'Histoire Naturelle, Paris)

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