Calosoma (Carabomorphus) catenatum Roeschke, 1899

catenatum and masaicum are both characterized by the disappearance of striation of elytra at the area of the basal pedicle, but catenatum is further characterized by the aedeagus with elongated apex, thin and curved. Instead, the inner armature of the penis is quite similar to masaicum as it has an elongated ligule with an apex laterally compressed and curved in the shape of crest.
Both species have some populations where the sculpture of the elytra has the intervals of the same width and height ("homodyname" sculpture) as well as other populations where the intervals on the elytra are unequal ("eterodiname" sculpture) , until you get to specimens with elytra smooth with only large foveae. In fact, the various populations of catenatum and masaicum are distinguishable only by examination of male genitalia.
Regarding catenatum, we can distinguish the population at relatively lower levels with the sculpture of the elytra quite regular (catenatum kenycola) from the population at higher altitudes with the sculpture of the elytra variously modified. (catenatum catenatum)

Calosoma (Carabomorphus) catenatum kenycola Jeannel, 1940

Carabomorphus kenycola Jeannel, 1940: 158 (type: prairies entre les rivières Amboni et Narèmuru; Muséum National d'Histoire Naturelle, Paris)
Carabomorphus catenatus ssp. meruensis Jeannel, 1940:159 (type: env. Meru, Mbagoris; British Museum of Natural History, London)
Carabomorphus catenatus ssp. kenycola Basilewsky, 1967:

Length 20-30 mm. It is found on the north-eastern and north-western slopes of Mount Kenya, at relatively medium altitude, around 2000-2800m.
Jeannel (1940: 158) described kenycola from the meadows between Mount Aberdare and Mount Kenya as a distinct species, and catenatus meruensis from a locality near Meru on the opposite western slopes of the same Mount Kenya. Both these populations have a sculpture with equal and regular intervals but on the unique specimen on which the description of meruensis is based, the primary intervals are fragmented into tubercles throughout their length.
Originally Jeannel attributed the two populations to two distinc species, on the base of slight differences of the form of penis and the apex of the ligule, but subsequently Basilewsky (1967) reviewed the question and considered also kenycola as a subspecies of catenatum .
So also in the case of catenatum seems to be verified the hypothesis that links the variation of the sculpture of elytra to the succession of the altitudinal belts.

Examined specimens
Kenya. Mt. Kenya, prairies entre les rivières Amboni et Narèmuru (holotype of kenycola, MNHN; RMCA); Mt. Kenya, Sirimon track, 2100 m (sub meruensis, RMCA)

Notes: Wingless. Catches are reported in December in the period immediately following the short rain.

Calosoma (Carabomorphus) catenatum kenycola
Jeannel, 1940
Kenya, Mt. Kenya, prairies entre les rivières Amboni et Narèmuru,
prateries découvertes 1900 2200m, Alluaud er Jeannel
(type of Carabomorphus kenycola);
(coll. Muséum National d'Histoire Naturelle, Paris)
Calosoma (Carabomorphus) catenatum kenycola
Jeannel, 1940
Kenya, Mt. Kenya, prairies entre les rivières Amboni et Narèmuru,
prateries découvertes 1900 2200m, Alluaud er Jeannel
(coll. Muséum National d'Histoire Naturelle, Paris)

Calosoma (Carabomorphus) catenatum kenycola
Jeannel, 1940
Kenya, Mt. Kenya (North), Sirimon track 2550m, 12.8.75, Dall’Asta
(sub catenatus meruensis)
(coll. Musée Royal de l'Afrique Central, Tervuren)
Calosoma (Carabomorphus) catenatum kenycola
Jeannel, 1940
Kenya, Mt. Kenya (North), Sirimon track 2550m, 12.8.75, Dall’Asta
(sub catenatus meruensis)
(coll. Musée Royal de l'Afrique Central, Tervuren)

Calosoma (Carabomorphus) catenatum catenatum Roeschke, 1899

Carabomorphus catenatus Roeschke, 1899:357 (typus: Kenya)
Carabomorphus catenatus ssp. Kolbi Roeschke, 1899: 358 (neotypus: Kenya; Musée Royal de l'Afrique Central, Tervuren)
Carabomorphus catenatus ssp. Bastinelleri Roeschke, 1899: 358
Calosoma (Carabomorphus) brachycerum ssp. catenatum Breuning, 1928: 135
Carabomorphus (Carabomorphus) catenatus Lapouge, 1932: 396
Carabomorphus catenatus ssp. jabbalae Basilewsky, 1967: (typus: Mt. Kenya; Musée Royal de l'Afrique Central, Tervuren)

Length 20-25 mm. The populations at higher altitudes (from 3000 up to 4100m) are of slightly reduced size, sometimes of squat forms and always with a more evoluted sculpture of the elytra.
In these populations the secondary intervals remain as continuous ribs, the tertiary ones are reduced or disappear altogether; the primary intervals are fragmented into large tubercles (catenatum), more or less confluent (kolbei). In some case, towards the upper limit where the species can live, also the primary intervals disappear, along with the secondary ones, and they only are indicated by series of large foveae (bastinelleri, jabbalae).

Examined specimens
Kenya. Mt. Kenya 3000-3300 m (AC; MNHN; neotype kolbi RMCA); Chogora Rt. 3000 m (PS); Kazità track 3350 m (sub bastinelleri, RMCA); Kazità West valley 3810 m (holotype jabbalae, RMCA)

Notes: Wingless. Catches are reported in July-August up to October-November, mostly during the wetter of the two dry seasons following the long rain.

Calosoma (Carabomorphus) catenatum catenatum
Roeschke 1899
(comp with type) unreadable label
(coll. Musée Royal de l'Afrique Central, Tervuren)
Calosoma (Carabomorphus) catenatum catenatum
Roeschke 1899
Mt. Kenya, Africa, XI.23, Arthur
(coll. Muséum National d'Histoire Naturelle, Paris)
Calosoma (Carabomorphus) catenatum catenatum
Roeschke 1899
Kenya: Mt. Kenya 3300m (coll. A. Casale)

Calosoma (Carabomorphus) catenatum catenatum
Roeschke 1899
Mt Kenya 10500ft, January, 1944, Mrs Bally
(neotype ssp. Kolbi Roeschke)
(coll. Musée Royal de l'Afrique Central, Tervuren)
Calosoma (Carabomorphus) catenatum catenatum
Roeschke 1899
Kenya: Mt. Kenya East 3000m, X.02,
W. Freund leg. (Museum König, Bonn)
Calosoma (Carabomorphus) catenatum catenatum
Roeschke 1899
Kenya: Mt. Kenya, Chogora Rt., 3000m,
4.X.01, Th. Wagner leg. (coll. P. Schüle)

Calosoma (Carabomorphus) catenatum catenatum
Roeschke 1899
Mt Kenya 11.000ft, Kazità track, IV 1966 Miss I. Jabbal
(sub Bastinelleri, Musée Royal de l'Afrique Central, Tervuren)
Calosoma (Carabomorphus) catenatum catenatum
Roeschke 1899
Mt Kenya 12.500ft, Kazità west valley, 28.IV 1966 Miss I. Jabbal
(holotype ssp. jabbalae Basilewsky)
(coll. Musée Royal de l'Afrique Central, Tervuren)

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