Calosoma (Callistenia) moniliatum LeConte, 1851

Calosoma moniliatum LeConte, 1851: 200 (described from Oregon & Washington; syntype in Museum of Comparative Zoology, Harvard University, Cambridge, Ma)
Calosoma laqueatum LeConte, 1851: 200 (described from Saskatshewan; syntype in Museum of Comparative Zoology, Harvard University, Cambridge, Ma)
Carabus bicolor Walker, 1866: 313 (nec Letzner 1850) (described from British Colombia; syntype in Natural History Museum, London)
Callisthenes concinnus Casey, 1913: 66 (described from Priest Lake, Idaho; lectotype in National Museum of Natural History, Washington)
Carabus taedatus var. vancouvericus Csiki, 1927: 286 (nomen novum pro bicolor Walker)
Calosoma (Callisthenes) moniliatum Breuning, 1928: 78
Calosoma (Callisthenes) moniliatum concinnum Breuning, 1928: 77
Callisthenes (Callistenia) moniliatus Lapouge, 1931: 378
Callisthenes (Callistenia) moniliatus var. concinna Lapouge, 1931: 378
Microcallisthenes (Callistenia) moniliatus Jeannel, 1940: 177
Callisthenes (Microcallisthenes) moniliatus Gidaspow, 1959: 305
Callisthenes (Callistenia) moniliatus Erwin, 2007: 79


Length 15-17 C. moniliatum, as C. wilkesi, can be easily sorted out by the elongated shape of the body. Both these two species have a transverse pronotum slightly narrower at the base, with well-conformed rear angles clearly protruding from the baseline.
The sculpture of the elytra of C. moniliatum is characterized by primary intervals with chain-like elevations interrupted by large foveae, these intervals have a glossy surface, while the remaining surface of the elytra is matt. The secondary intervals consist in series of aligned grains and the tertiary ones are transformed in a confuse granulation. The color of the upper body is dark bronze, almost blackish in some cases, in other cases the color can be clear and bright.
C. moniliatum is relatively common in the mountainous areas of southern Canada and United States, on the western slopes of the Rocky Mountains and of Sierra Nevada.

Examined specimens and literature’s data
Canada. Alberta: Waterton Lakes National Park, Waghorn, Magrath, Happy Valley, Millarville, Didsbury, Olds, Bowden, Blackfalds (Lindroth, 1961: 53), Jumpingpound Creek, Bragg Creek, Prairie Bluff (UASM), Calgary (AC); British Columbia: Waneta, East Kootenay, Radium (UASM), Vancouver (EM), Cranbrook, Creston (Lindroth, 1961: 53); Saskatchewan: Sylvan Lake NP, Saskatoon (Lindroth, 1961: 53), Baldwinton (UASM); Nova Scotia (http://carabidae.org/)
United States. Arizona, California (Gidaspow, 1959: 305); Idaho: Priest Lake (lectotype of concinnus), Idaho city (Breuning, 1928b: 77), Coolin (www.gbif.org); Nebraska, Montana, Oregon (Gidaspow, 1959: 305); Washington: Spokane (Breuning, 1928b: 77).

Notes: Contrary to the adjective “brachypterous” used by Jeannel (1940: 177), the wings of C. moniliatum are normally formed but, because of some peculiarities of their structure, Lindroth (1961: 51) expresses doubts about its ability to fly. C. moniliatum is found in uplands, from 1200 to 2200m altitude, on praries and dry forests. It is preferably nocturnal, taking refuge under stones in daytime (Erwin, 2007: 79). Adult are active mostly in early spring up to August.

Calosoma (Callistenia) moniliatum
LeConte, 1851
Canada: Alberta, Calgary, 1911 (coll. A. Casale)
Calosoma (Callistenia) moniliatum
LeConte, 1851
Canada: British Columbia, Vancouver (coll Migliaccio)

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